Crustacean

Etymology and History of Study

The term crustacean derives from the Latin word crusta, meaning a hard shell, crust, or rind, reflecting the rigid exoskeleton that covers the body.^[1][3][5] Early European naturalists, including Pierre Belon in 1551 and 1555 and Guillaume Rondelet in 1554 and 1555, employed the designation in their pioneering natural history treatises.^[5] In his tenth edition of Systema Naturae published in 1758, Carl Linnaeus omitted the group name Crustacea, instead subsuming these animals alongside other wingless arthropods under the order Aptera.^[6][7][8] The earliest taxonomically valid introduction of the Latinized subphylum name Crustacea was established by the Danish zoologist Morten Thrane Brünnich in his 1772 work Zoologiae Fundamenta, although Brünnich originally included chelicerates in the assemblage.^[6][7][8] Brünnich was later followed by Thomas Pennant in 1777, Georges Cuvier in 1799 to 1800, and Jean-Baptiste Lamarck in 1801..^[7]

Aristotle initially documented decapods and their allies under the Greek designation malakostraka, distinguishing them from terrestrial insects and classifying seventeen marine species.^[7][9] In the eighteenth and nineteenth centuries, scholarship progressed through contributions by Louis Jean-Marie Daubenton in the Encyclopedie, Lamarck in his natural history of invertebrates, Constantine Samuel Rafinesque, Charles Darwin in his monographs on cirripedes, and Henri Milne Edwards in his multi-volume Histoire naturelle des Crustaces, which named hundreds of genera and species.^[10] The scientific discipline dedicated to the study of crustaceans is known as carcinology, with malacostracology, crustaceology, and crustalogy occasionally serving as alternative terms, and its practitioners are termed carcinologists.^[11]

External Anatomy and Morphology

The crustacean body varies widely in architectural organization, ranging from simple, elongated, multi-segmented trunks to heavily compact, fused bodies.^{[3][12][13][14]} Overall body dimensions span from microscopic parasites like Stygotantulus stocki, measuring less than 0.1 millimetres (0.004 inches) in total body length, and Tantulacus dieteri at only 85 micrometres, to the Japanese spider crab (Macrocheira kaempferi), which can attain a leg span of 3.8 metres (12.5 feet) and a mass of 20 kilograms (44 pounds).^{[3][15][16][17]} The American lobster (Homarus americanus) is the heaviest crustacean and can weigh more than 20 kilograms, while the giant Tasmanian crab (Pseudocarcinus gigas) reaches up to 14 kilograms..^[18] On land, the coconut crab (Birgus latro) represents the largest terrestrial invertebrate, weighing up to 4 kilograms.^[18]

The basic body plan comprises a chain of body segments, known as somites or metameres, delimited anteriorly by an unsegmented pre-oral acron and posteriorly by a terminal telson.^[3][12][19] Primitive or ancestral conditions exhibit homonomous segmentation with as many as fifty or more body segments, seen in certain branchiopods and remipedes.^{[3][14][19][20]} More derived groups display heteronomous tagmosis, consolidating metameres into distinct functional tagmata: the head or cephalon, the thorax or pereon, and the abdomen or pleon.^{[3][12][19][21][22]} Decapods and many other malacostracans typically possess a fixed number of segments: the cephalon consists of the acron plus five metameres, followed by an eight-segmented thorax and a six-segmented abdomen ending in the telson, whereas leptostracans uniquely retain seven abdominal segments.^[4][19]

Tagmata, Carapace, and Sclerites

The cephalon comprises the acron and five more or less fused segments, which bear the antennules, antennae, mandibles, maxillules, and maxillae..^{[3][12][19][23]} In many lineages, the cephalon is divided or differentiated into an anterior protocephalon, formed by the acron and antennular segment, and a posterior gnathocephalon, bearing the feeding appendages.^[24][25] Frequently, one or more thoracic segments fuse with the head to produce a cephalothorax.^[19][26][27] The dorsal integument of the head often expands backward as a fold of cuticular tissue called the carapace.^{[3][19][28][29]} The carapace varies from an unobtrusive cephalic shield to a structure covering the thorax, as in lobsters and crayfish, or a bivalved mantle enveloping the entire body, as seen in ostracods, clam shrimp, and ascothoracids.^{[3][19][29][30]} In cirripedes, this mantle secretes calcareous plates that form a rigid, enclosing shell.^[31]

The exoskeleton of each segment consists of cuticular plates or sclerites.^[3][32] The dorsal plate is termed the tergum or tergite, the ventral plate is the sternum or sternite, and the lateral connecting regions are the pleura or pleurites.^{[3][19][29][32]} The cuticular exoskeleton is secreted by a single layer of hypodermal cells and contains chitin bonded with structural proteins.^[3][33][34] In larger crustaceans, the procuticle is extensively mineralised by deposits of calcium carbonate and calcium phosphate, giving the shell its characteristic rigidity.^[3][33][34] Flexible, non-calcified membranes remain at joints between segments and podomeres, enabling articulation and limb locomotion.^[3]

Appendage Architecture and Specialisation

Crustacean appendages are fundamentally biramous, consisting of a basal stalk termed the protopodite, which typically comprises two podomeres: the coxa or coxopodite and the basis or basipodite.^{[3][14][19][35]} In some taxa, a proximal precoxa is integrated into the body wall.^[36] Distally, the basis articulates with two branches: an outer exopodite and an inner endopodite.^[3][14][19] The protopodite may bear lateral cuticular projections called exites, which frequently function as respiratory epipodites, as well as medial projections termed endites.^[3][14][19] The endites of feeding limbs often develop heavily sclerotised, dentate chewing processes known as gnathobases.^[3][14][19]

Appendages diverge into two broad structural classes: stenopodia and phyllopodia.^[37][38] Stenopodia are narrow, tubular, strongly jointed limbs with thick cuticular walls, adapted primarily for walking, grasping, and burrowing, such as the walking legs of decapods.^[37][38] In typical malacostracan walking legs, the endopodite comprises five podomeres: the ischium (ischiopodite), merus (meropodite), carpus (carpopodite), propodus (propodite), and dactylus (dactylopodite).^[36] Phyllopodia are broad, flattened, leaf-like limbs with thin cuticles and expansive epipodites, serving simultaneously for swimming, suspension feeding, and gas exchange, as observed in branchiopods, cephalocarids, and leptostracans.^[37][38]

The cephalic tagma bears five pairs of appendages: the first antennae or antennules, the second antennae, the mandibles, the first maxillae or maxillules, and the second maxillae.^{[3][12][14][19]} Antennules are uniramous in most crustaceans, though malacostracans frequently possess biramous or triramous antennules.^[19][35] Copepods often move through the water using their first pair of antennae.^[39] In some non-parasitic species, males have enlarged antennules that hold the female during copulation..^[40] The second antennae are biramous in ancestral forms, with their exopodites sometimes flattened into a scaphocerite or antennal scale.^[14][19] The mouth opening is bordered anteriorly by the labrum and posteriorly by the paragnaths, while the heavily sclerotised mandibles crush food items.^[3][24][41]

Thoracic appendages, or thoracopods, are attached to the pereon.^{[3][19][21][36]} When anterior thoracic segments fuse with the head, their appendages are modified into feeding accessories known as maxillipeds, or foot-jaws, which hold and manipulate food toward the mouth; decapods possess three pairs of maxillipeds.^{[3][19][21][42]} Unmodified thoracic limbs are called pereiopods, commonly functioning as walking legs.^{[3][19][21][42]} In astacideans and brachyurans, the first pair of pereiopods is often enlarged into massive chelipeds bearing opposable claws or chelae, while true crabs possess a single pair of chelae and penaeid shrimps possess three pairs of smaller chelae.^[16][42]

Abdominal appendages, or pleopods, are present in malacostracans and remipedes, but are absent or vestigial in other crustacean classes.^{[3][19][42][43]} Pleopods typically serve swimming functions, maintain respiratory water currents, or carry developing egg broods in females.^[3][19][42] In male decapods, the first and second pairs of pleopods are modified into sclerotised intromittent gonopods for transferring spermatophores.^[42][44][45] The terminal abdominal segment is the telson, which bears the anus and may carry paired caudal rami forming a caudal furca.^{[3][19][43][46]} In many malacostracans, the sixth pair of pleopods is transformed into broad, flattened uropods that flank the telson to form a tail fan, utilized in rapid backward escape swimming.^{[3][19][42][46]}

Internal Anatomy and Physiology

The crustacean coelom is reduced during embryonic development to remnants surrounding the gonadal cavities and the end-sacs of the excretory organs.^[40][47] The primary body cavity is a haemocoel, through which haemolymph circulates in an open circulatory system driven by a dorsal heart.^{[3][48][49][50]} The heart resides within a pericardial sinus and draws in haemolymph via paired lateral slits called ostia.^[50][51][52] The structural complexity of the heart varies extensively: primitive branchiopods and remipedes possess an elongate tubular heart extending along the dorsal trunk with segmental ostia, while cladocerans and decapods have a compact, muscular, vesicle-like heart.^[50][51] In decapod crustaceans, the heart pumps blood into arterial vessels that branch into capillary-like networks before discharging into tissue sinuses.^[40][50][53]

Respiratory pigments in the haemolymph facilitate gas transport. Malacostracans employ copper-based haemocyanin dissolved freely in the blood plasma, whereas copepods, ostracods, barnacles, and branchiopods rely on haemoglobins.^{[40][49][54][55]} Some malacostracans additionally maintain tissue haemoglobins.^[40] In cyclopoid copepods the circulatory structures disappear entirely.^[56] Some groups within Ostracoda, Copepoda, and Cirripedia lack well-defined vessels..^[56][57]

Alimentary System and Digestion

The digestive tract of crustaceans is a straight or slightly curved tube composed of an ectodermal foregut, an endodermal midgut, and an ectodermal hindgut.^{[3][29][58][59]} The foregut and hindgut are lined with chitinous cuticle that is shed and replaced during each moult.^[59][60] The foregut consists of a short muscular oesophagus and a stomach.^[29][59][61] In malacostracans, the stomach is differentiated into two chambers: a voluminous anterior cardiac stomach and a smaller posterior pyloric stomach, separated by internal valves.^[29][59][61] The cardiac stomach houses the gastric mill, an internal masticatory apparatus consisting of calcified cuticular ossicles that form one dorsal and two lateral grinding teeth operated by extrinsic muscles.^{[29][58][59][61]}

The pyloric stomach functions as a sorting chamber, using parallel rows of setae to filter finely ground particles from coarse, indigestible debris.^[29][59][61] Fine particles and liquid nutrient fractions enter the midgut, which is connected to large branched digestive diverticula termed the hepatopancreas or midgut gland.^{[58][61][62][63]} The hepatopancreas consists of blind-ending tubules lined with specialized epithelial cells that synthesize and secrete digestive enzymes, execute intracellular and extracellular digestion, absorb nutrients, and store glycogen, lipids, and calcium salts.^[61][62][63] Coarse indigestible particles are directed into the midgut lumen, encapsulated by a thin, transparent peritrophic membrane secreted by the midgut epithelium, and conveyed through the cuticular hindgut to be voided via the anus on the telson.^[61][64]

Respiration, Excretion, and Osmoregulation

Gas exchange in crustaceans occurs either across the general body surface or through specialized respiratory organs.^[40][49][65] Very small crustaceans, including many copepods and ostracods, possess high surface-area-to-volume ratios and thin cuticles, allowing direct cutaneous diffusion of oxygen and carbon dioxide.^[40][49][50] Larger aquatic crustaceans rely on gills, which represent modified epipodites of thoracic limbs or branchial outgrowths of the body wall.^{[40][49][65][66]} In decapods, gills are enclosed within bilateral branchial chambers sheltered under the lateral expansions of the carapace, termed branchiostegites.^[36][40][49] Water is actively pumped through these chambers by the rhythmic beating of the scaphognathite, or gill bailer, an exopodite of the second maxilla capable of beating up to 200 times per minute.^[40][49][65]

Terrestrial adaptations have prompted modifications to respiratory structures. Land isopods (woodlice) possess pleopodal lungs, or pseudotracheae, which are branched internal invaginations on the exopodites of their abdominal pleopods that permit aerial respiration while minimizing desiccation.^{[40][49][50][67]} In terrestrial crabs, the gills are frequently reduced in size, and the inner lining of the branchiostegite is vascularized and transformed into a lung chamber, while thin membranous cuticular discs on the legs facilitate accessory gas exchange in ghost crabs.^[40][49][68]

Excretion and osmoregulation are managed primarily by paired segmental nephridia, which open either at the bases of the second antennae as antennal glands (also called green glands) or at the bases of the second maxillae as maxillary glands.^{[40][69][70][71]} Adult malacostracans generally have antennal glands, whereas other crustaceans generally have maxillary glands.^[70][71] Both pairs occur in Nebalia and marine ostracods, and also in Mysida and Lophogastrida..^[40][70][72] Each excretory gland begins as a closed coelomic end-sac (sacculus) lined with podocytes, where ultrafiltration of haemolymph occurs under hydrostatic pressure.^{[40][70][71][73]} The filtrate passes into an extensive, coiled tubule or spongy labyrinth that selectively reabsorbs essential ions, glucose, and water before conveying the fluid to a distal bladder that empties through an excretory pore.^{[40][70][71][73]}

Freshwater crustaceans maintain elongated nephridial tubules to produce dilute, hyposmotic urine, thereby conserving salts, whereas marine species maintain shorter tubules producing isosmotic urine.^[73] Crustaceans are predominantly ammonotelic, eliminating over 90 percent of their nitrogenous waste as toxic ammonia.^[40][74] Much of this ammonia is discharged across the thin branchial epithelium rather than via the nephridiopores.^[40][70] In terrestrial isopods, small quantities of uric acid are excreted in addition to gaseous ammonia.^[40] Additional excretory functions are performed by phagocytic nephrocytes scattered through the haemocoel and the bases of the legs, which sequester and break down particulate metabolic wastes.^[40][70]

Nervous, Sensory, and Endocrine Systems

The central nervous system follows the ancestral arthropod ladder-like plan, consisting of a dorsal tripartite brain linked via paired circumoesophageal connectives to a suboesophageal ganglion and a ventral nerve cord with paired segmental ganglia.^[75][76][77] The brain comprises the protocerebrum, which innervates the eyes and incorporates associative hemiellipsoid bodies analogous to insect mushroom bodies; the deutocerebrum, which innervates the antennules; and the tritocerebrum, which innervates the antennae and connects to the visceral nervous system.^{[75][76][78][79][80]} Extensive ganglionic fusion has evolved independently in multiple lineages: crayfish show midline fusion of segmental ganglia into a single ventral nerve cord, while brachyuran crabs have concentrated all thoracic and abdominal ganglia into a single massive ventral thoracic nerve mass.^[75][76][81]

Visual reception is mediated by both median simple eyes and lateral compound eyes.^[75][76][82] The larval nauplius eye consists of two to four inverted pigment-cup ocelli.^[76][83] It is retained in adults of cyclopoid copepods and Triops, but is typically superseded by lateral compound eyes in adult malacostracans.^[75][76][82] Compound eyes consist of numerous visual subunits called ommatidia, each possessing a corneal lens, crystalline cone cells, retinular cells, and a central rhabdom.^[84][85][86] In mantis shrimp (Stomatopoda), the compound eyes reach extraordinary complexity, possessing up to twelve photoreceptor channels for color discrimination across ultraviolet and visible wavelengths, alongside dedicated channels for linear and circular polarized light.^[87] Blindness and eye loss occur frequently in subterranean, cave-dwelling, and deep-sea taxa.^[75][76][82]

Cuticular sensory organs include innervated setae functioning as tactile mechanoreceptors and aesthetascs acting as chemoreceptors on the antennules.^[76][86][88] Statocysts, serving as equilibrium organs, are positioned at the base of the antennules in decapods and in the uropodal endopodites of mysids.^[75][86] Decapod statocysts consist of fluid-filled invaginations containing sensory setae and sand grains, which act as statoliths and must be replaced from the substrate after every moult.^{[75][76][86][89]} Spiny lobsters can navigate using geomagnetic fields, while Cherax crayfish demonstrate electroreceptive capabilities.^[90]

Endocrine regulation is integrated with neurosecretory centres, predominantly the X-organ/sinus-gland complex located within the eyestalks of stalk-eyed decapods.^{[91][92][93][94]} The X-organ produces neurohormones stored and released by the sinus gland, notably moult-inhibiting hormone (MIH), vitellogenesis-inhibiting hormone (VIH), crustacean hyperglycaemic hormone (CHH), and chromatophorotropins.^[91][92][93] Crustecdysone, the active ecdysteroid hormone that stimulates premoult processes and ecdysis, is synthesized by the paired Y-organs situated in the cephalothorax, whose activity is normally restrained by MIH.^{[92][93][94][95]} Decapod sexual differentiation is governed by the androgenic gland, located along the vas deferens, which secretes hormones directing the differentiation of male reproductive morphology and secondary sexual traits.^[92]

Pigmentation and physiological colour change are driven by chromatophores in the hypodermis.^[29][96] Chromatophores contain black, red, yellow, or white pigment granules that disperse or concentrate in response to pigment-dispersing hormones (PDH) and pigment-concentrating hormones (PCH).^[96][97] Heart rate and peripheral muscular tone are modulated by cardioactive amines, including octopamine, dopamine, and serotonin (5-hydroxytryptamine), released into the pericardial space by the pericardial organs.^[92][98] Serotonin elevates motor neuron transmission and aggression, promoting combat behavior in crayfish, while octopamine fosters submissive behaviors.^[98][99]

Life Cycle, Reproduction, and Development

Most crustaceans are gonochoric, possessing separate sexes and reproducing through sexual copulation, though diverse alternative modes exist.^{[3][100][101][102]} Simultaneous hermaphroditism is characteristic of Cephalocarida, Remipedia, and thoracic barnacles, where sessile adults benefit from cross-fertilization with neighboring individuals through an extremely elongate, extensible penis.^{[40][101][103][104][105]} Sequential hermaphroditism occurs in several decapod groups, including protandric northern shrimp (Pandalus borealis).^[36] Parthenogenesis occurs in certain branchiopods, ostracods, and isopods, as well as the marbled crayfish (Procambarus virginalis), which reproduces obligately without males.^{[3][100][101][102]}

Crustacean sperm are typically non-motile and lack flagella.^[61][106] Flagellated sperm occur in remipedes, cirripedes, mystacocarids, and ostracods..^[61] Ostracods have flagellated, motile sperm, and in some species part of the male sperm duct forms a Zenker organ that works as a peristaltic sperm pump.^[61] Propontocypris monstrosa measures about 0.6 millimetres, but its sperm are 5 to 7 millimetres long.^[107] Transfer of such large sperm depends on special muscular organs in the female reproductive tract..^[107][108] Male crustaceans transfer sperm packaged within protective spermatophores, using modified appendages such as the petasma in penaeid prawns or the first and second pleopods in higher decapods.^{[45][109][110]} Females often store sperm in an internal seminal receptacle or an external spermatheca until ovulation.^{[40][45][110]}

Parental brooding of eggs is widespread across Crustacea.^{[3][102][111]} Decapod females attach fertilized eggs to their abdominal pleopods with adhesive secretions, aerating them until hatching.^{[3][100][102][111]} Peracarids develop a ventral brood pouch, or marsupium, enclosed by overlapping thin plates called oostegites originating from the thoracic coxae, where young undergo direct development.^{[45][100][102]} Cladocerans and thermosbaenaceans incubate embryos within a dorsal brood chamber formed beneath the carapace.^{[45][61][100][102]} Branchiuran fish lice glue rows of fertilized eggs to submerged rocks.^[112][113] In adverse conditions, branchiopods produce thick-shelled resting eggs or cysts, such as the ephippia of Daphnia, which endure desiccation and freezing for years.^{[61][114][115]}

Development proceeds via anamorphic, metamorphic, or epimorphic trajectories.^{[3][116][117]} The ancestral free-swimming larva is the nauplius, which possesses an unsegmented body, a median naupliar eye, and three pairs of appendages: uniramous antennules and biramous antennae and mandibles, which provide propulsion.^{[3][117][118][119]} Subsequent moults add post-mandibular segments in an anamorphic metanauplius stage.^{[3][117][120]} Advanced metamorphic phases appear in malacostracans, notably the zoea larva, which possesses a cephalothorax, compound eyes, thoracic swimming appendages, and a segmented abdomen.^{[3][116][117][118][121]} Decapod larvae often pass into a mysis, megalopa, or phyllosoma stage before settling.^{[116][117][121]} In many freshwater decapods, including astacid crayfish and potamid crabs, larval stages are suppressed in favor of direct epimorphic development, hatching as miniature juveniles.^{[3][116][122]}

Growth requires periodic moulting, or ecdysis, of the cuticular exoskeleton, divided into proecdysis (premoult), ecdysis, metecdysis (postmoult), and aneuroecdysis (intermoult).^{[92][123][124]} In proecdysis, hypodermal enzymes dissolve and resorb calcium and organic compounds from the inner endocuticle into the haemolymph, while a flexible new epicuticle and exocuticle are synthesized underneath.^[124] The old exoskeleton ruptures along predefined ecdysial suture lines, and the animal withdraws its body, rapidly inflating its volume through water absorption before hardening the new cuticle by mineralizing calcium carbonate.^[124] Research on DNA repair in black tiger shrimp (Penaeus monodon) indicates that double-strand breaks are repaired predominantly by accurate homologous recombination and microhomology-mediated end joining.^[125] Ultraviolet irradiation in Tigriopus japonicus copepods triggers increased expression of proteins involved in non-homologous end joining, base excision repair, and mismatch repair.^[126]

Feeding Mechanisms and Trophic Modes

Crustaceans exploit virtually every trophic niche, including suspension feeding, deposit feeding, active predation, scavenging, herbivory, and parasitism.^{[37][127][128]} Suspension feeders capture suspended phytoplankton, bacteria, and fine organic particles using specialized setae.^[3][129] Anostracan and cladoceran branchiopods generate feeding currents with their phyllopodal limbs, drawing water into suction chambers between limb bases where fine setae filter particles toward a ventral food groove leading to the mouth.^{[3][129][130]} In adult cirripedes, feeding is conducted by extending and retracting the six pairs of multi-articulated, plumose thoracic cirri, which form a sweep-net to catch microplankton.^{[129][131][132]} Copepods have well-developed chemoreceptors, especially on their mouthparts and feeding appendages, which let them detect phytoplankton cells in the water and capture them with asymmetric movements of the appendages..^[133]

Among malacostracans, Antarctic krill (Euphausia superba) form a thoracic feeding basket using elongate, setose pereiopods, compressing water outward while trapping diatoms and microzooplankton.^[129][134] Porcelain crabs (Porcellanidae) employ fan-shaped setae on their third maxillipeds to filter suspended matter, switching from active sweeping in still water to passive posture in strong currents.^[135][136] Deposit feeders harvest benthic detritus, organic coatings, and meiofauna from soft sediments.^[3][129] Fiddler crabs (Uca) and sand-bubbler crabs (Scopimera) gather surface substrate with their chelae, sorting organic particles using spoon-tipped maxilliped setae while rejecting mineral sand as discarded pellets.^{[3][129][137][138]} Burrowing mud shrimps (Thalassinidea) extract food by filtering pumped burrow currents or scraping mud from tunnel walls.^[139]

Predators and scavengers comprise a substantial proportion of malacostracan diversity.^[3][128] Many crabs break the shells of thick-shelled molluscs with heavy claws, whose muscles can produce crushing forces of up to 800 newtons..^{[140][141][142]} Mantis shrimps (Stomatopoda) utilize raptorial thoracic limbs that strike prey with speeds of two milliseconds, delivering blows comparable to small-caliber projectiles that fracture snail shells and crab carapaces.^{[143][144][145]} Pistol shrimps of the genus Alpheus snap an enlarged asymmetrical claw shut, producing high-velocity cavitation bubbles that emit intense shock waves to stun or kill prey.^[129] Remipedes represent the only known venomous crustaceans; Xibalbanus tulumensis delivers a complex toxic cocktail containing neurotoxic peptides, chitinases, and xibalbins via modified prehensile maxillules to paralyze prey.^[146][147]

Deep-sea lysianassoid amphipods and isopods (such as Bathynomus) function as opportunist scavengers on benthic carcasses, possessing specialized cutting mouthparts and distensible guts that allow rapid engorgement on vertebrate carrion.^{[3][148][149]} Detritivores, including terrestrial isopods and land crabs like Gecarcoidea natalis, feed on vascular leaf litter and wood debris, processing cellulose with mechanical gastric mills and intestinal microbiota.^{[3][150][151][152]} Certain species, such as the freshwater crayfish Cherax destructor, possess endogenous cellulases, allowing them to digest plant structural carbohydrates without relying entirely on microbial symbionts.^[153] Coprophagy is practiced by terrestrial isopods to reingest nutrient-rich, microbially conditioned faeces, functioning as an external rumen.^[3]

Symbiosis and Parasitism

Parasitism has arisen independently in multiple crustacean lineages, often resulting in profound reductions and modifications of standard arthropod anatomy.^{[40][154][155]} The Ichthyostraca comprises branchiurans and pentastomids.^[156] Branchiurans (fish lice, such as Argulus) are temporary ectoparasites on fish skin and gills, utilizing modified sucking maxillae and piercing stylets.^[157] Pentastomids (tongue worms) are endoparasites residing in the respiratory tracts of reptiles, birds, and mammals, possessing unsegmented vermiform bodies with four hook-like claws around the mouth.^[158][159] Within Thecostraca, rhizocephalan barnacles, such as Sacculina, parasitize decapod crabs.^[154][160] A female cyprid larva settles on a crab, sheds its extremities, and injects an unsegmented vermigon into the host haemocoel; this develops into an extensive, nutrient-absorbing root network (interna) that eventually pushes an external reproductive sac (externa) out under the crab abdomen, sterilizing and hormonally feminizing male hosts to groom and care for the parasite brood.^[154][160]

Tantulocarids are microscopic ectoparasites of deep-sea crustaceans, with an adhesive cephalic disc that penetrates the host cuticle.^{[40][161][162]} Copepods include ectoparasitic caligids (sea lice) and highly transformed mesoparasitic forms like Lernaeocera and Pennella, the latter reaching lengths of thirty centimetres on marine mammals.^{[40][154][160][163]} Among isopods, bopyrids parasitize decapod branchial chambers, and Cymothoa exigua parasitizes marine fish, clinging to and consuming the blood supply of the host tongue until the organ atrophies, whereupon the isopod attaches to the remaining muscular stub and functionally replaces the tongue.^{[61][164][165]} Conversely, many cleaner shrimps (Palaemonidae and Hippolytidae) engage in mutualistic cleaning symbioses, removing ectoparasites from reef fishes.^[166] Eusocial colonies occur in Synalpheus sponge-dwelling shrimps, where reproductive queens and kings are defended by sterile snapping workers.^[167]

Habitats, Ecology, and Environmental Roles

Crustaceans dominate marine biomes across all bathymetric zones, occupying epipelagic waters, abyssal plains, benthic substrates, hydrothermal vents, and hadal trenches below 9,000 metres.^{[3][18][85][168]} Copepods are the dominant component of marine zooplankton from cold regions to the tropics, and marine planktonic crustaceans, largely copepods, take part in daily vertical migrations, rising through the water column at night to feed and sinking to greater depths by day to avoid predators.^[169][170] In the Southern Ocean, the Antarctic krill (Euphausia superba) reaches an estimated biomass of roughly 500 million tonnes and a population of 700 trillion individuals, forming the keystone prey resource sustaining fishes, penguins, seals, and baleen whales.^{[5][18][171][172]}

Freshwater environments support diverse assemblages of branchiopods, amphipods, isopods, and decapods.^{[3][173][174]} Anostracans, notostracans, and clam shrimps thrive in ephemeral rain pools, vernal ponds, and hypersaline lakes like the Great Salt Lake, where Artemia salina tolerates extreme osmotic pressures.^{[61][173][174]} Subterranean and stygobitic habitats harbor specialized eyeless, unpigmented taxa, such as bathynellaceans, first discovered in a Prague well by Frantisek Vejdovsky, and thermosbaenaceans, originally described from hot springs in Tunisia.^{[173][175][176]} Anchialine limestone caves shelter ancient remipedes that swim upside down in stratified brackish groundwater.^[177]

Terrestrial habitats have been successfully colonized by multiple lineages.^{[3][151][178]} Oniscidean isopods (woodlice) encompass over 5,000 terrestrial species found from humid forest leaf litter to arid deserts, such as Hemilepistus reaumuri in North African sand environments, and high-altitude slopes exceeding 4,700 metres.^[25][151] Terrestrial decapods include land hermit crabs (Coenobita), ghost crabs (Ocypode), and land crabs (Gecarcinidae), which can process substantial volumes of forest litter on tropical islands.^[3][151] On Christmas Island, tens of millions of Gecarcoidea natalis undertake synchronized annual breeding migrations from inland rainforests to the ocean shore to deposit eggs into seawater.^[44][179]

Several crustacean species act as ecosystem engineers and invasive pests.^{[180][181][182][183]} Thalassinidean burrowing shrimps disrupt marine sediments through extensive bioturbation, altering benthic nutrient flux and community structures.^[182] Sphaeromatid isopods burrow into red mangrove aerial prop roots, altering root architecture.^[129] Invasive crustaceans include the Chinese mitten crab (Eriocheir sinensis), which was brought to Europe, where it erodes riverbanks and tears fishing nets; the Asian shore crab (Hemigrapsus sanguineus); the American spinycheek crayfish (Faxonius limosus); and the killer shrimp (Dikerogammarus villosus).^{[180][181][183][184]} Since the 1869 opening of the Suez Canal, nearly 100 Indo-Pacific and Red Sea crustacean species have established populations in the eastern Mediterranean Sea via Lessepsian migration.^[185]

Classification and Phylogeny

Historically, crustaceans were organized morphologically into two broad categories: lower crustaceans (Entomostraca) and higher crustaceans (Malacostraca).^[186][187] Bowman and Abele in 1982 codified a system of six living classes: Branchiopoda, Remipedia, Cephalocarida, Maxillopoda, Ostracoda, and Malacostraca.^[188] In 2001, Joel W. Martin and George E. Davis updated this framework to encompass 849 extant families across 42 orders, maintaining the six classes while noting that Maxillopoda was non-monophyletic.^[189][190] Morphological groupings also posited either a Malacostraca-Entomostraca sister relationship or a Maxillopoda-Thoracopoda grouping.^[191][192]

Molecular phylogenetics and cladistic analyses demolished the monophyly of both Maxillopoda and traditional Crustacea.^{[193][194][195][196][197][198][199]} Nuclear and mitochondrial gene sequencing, transcriptomics, and neuroanatomical studies demonstrated that Hexapoda (insects, springtails, proturans, and diplurans) originated from within the crustacean tree.^{[2][193][197][200][201]} Consequently, Crustacea is paraphyletic unless defined to include hexapods under the monophyletic clade Pancrustacea (also named Tetraconata due to the shared presence of four crystalline cone cells in each ommatidium).^{[2][201][202][203][204][205]} Maxillopoda was abandoned as a polyphyletic assemblage, and its subclasses were elevated to class or superclass ranks.^{[196][206][207][208]}

In the system used by the World Register of Marine Species, living crustaceans are placed in three superclasses: Oligostraca, Multicrustacea, and Allotriocarida..^{[4][209][210][211]} Oligostraca forms the basal clade, uniting Ostracoda (seed shrimps), Mystacocarida, and Ichthyostraca (containing the parasitic Branchiura and Pentastomida).^{[4][199][210][212][213]} The remaining taxa constitute Altocrustacea, which branches into Multicrustacea and Allotriocarida.^{[4][210][214]} Multicrustacea comprises Malacostraca and Hexanauplia (which unites Copepoda, Thecostraca, and Tantulocarida), though some phylogenomic analyses suggest copepods may be sister to or nested within Allotriocarida.^{[210][214][215][216][217]} Allotriocarida unites Cephalocarida, Branchiopoda, Remipedia, and Hexapoda.^{[4][210][218][219]} Remipedia and Hexapoda form the well-supported clade Labiocarida.^[4][219] Morphological traits supporting it include a labium, neuroanatomy, and the shared presence of haemocyanin..^{[4][220][221]}

Recognised Classes and Groups

Under updated classifications, extant crustacean lineages represent ten to twelve distinct classes or sublineages:^{[4][206][207][222]}

Branchiopoda encompasses fairy shrimp (Anostraca), tadpole shrimp (Notostraca), clam shrimp (Laevicaudata, Spinicaudata, and Cyclestherida), and water fleas (Cladocera), largely occurring in fresh and temporary waters.^[4] Cephalocarida lack compound eyes.^[76] They live on the sea floor and come from muddy bottoms rich in organic material..^[223] Remipedia comprises blind, elongate, venomous cave-dwelling crustaceans swimming in anchialine environments.^[177] Ostracoda contains thousands of seed shrimp species possessing bivalved calcareous carapaces.^[4]

Mystacocarida are so small that they live between sand grains..^[224] Ichthyostraca unites the ectoparasitic Branchiura (fish lice) and the endoparasitic Pentastomida (tongue worms).^[4] Copepoda includes free-living planktonic and benthic species as well as parasitic forms.^[163][173] Copepods lack compound eyes, though they can have complex naupliar eyes..^[75][225] Tantulocarida are parasites 0.15 to 0.3 millimetres long and are among the smallest crustaceans, highly specialised for parasitic life..^{[4][40][162][226]} Thecostraca comprises sessile barnacles (Cirripedia), ascothoracids, and enigmatic facetotectans.^[4] Malacostraca is the largest class, comprising mantis shrimp (Stomatopoda), krill (Euphausiacea), isopods, amphipods, cumaceans, tanaids, and decapods.^[4]

Fossil Record and Geological History

Crustaceans have an extensive fossil record extending back to the early Cambrian period, approximately 510 to 520 million years ago.^{[227][228][229][230]} The earliest definitive fossils include Canadaspis and Perspicaris from the middle Cambrian Burgess Shale, alongside miniature, three-dimensionally preserved microfossils from the Swedish Orsten lagerstatte.^{[229][231][232][233]} Orsten deposits yield phosphatized larval and juvenile stages displaying precise cuticular details, establishing that early Cambrian stem-crustaceans possessed compound eyes, naupliar limbs, and typical mouthparts.^[229][232] Phosphatocopines, bivalved arthropods from the upper Cambrian, are regarded as stem-taxa closely allied to crown crustaceans.^[231][234]

Most major crustacean classes were established before the close of the Cambrian, including branchiopods, malacostracans, thecostracans, and early larval pentastomids.^[227][235] Ostracods, whose heavily calcified bivalved shells favour fossilization, have a continuous fossil record; Cambrian animals have been assigned to them, but it is debated whether these are true ostracods, which would otherwise first appear in the Ordovician.^[61][227] Fossil ostracods are commonly used as indicators of the age of sediments and of past climates.^[236] Freshwater branchiopods, including Lepidocaris rhyniensis (order Lipostraca), are preserved in the Devonian Rhynie Chert.^[237][238] Remipedes first appear as Tesnusocaris goldichi in Carboniferous strata, during which fossil crustaceans become globally abundant.^{[239][240][241]}

Within Malacostraca, isopods, cumaceans, and true mantis shrimp are recognized from Carboniferous deposits.^{[242][243][244][245]} Ghost shrimps (Axiidea, Gebiidea) and freshwater crayfish (Astacidea, Parastacidea) left distinct fossil burrows (Ophiomorpha and Camborygma) dating from the Permian Roadian stage in Nurra, Sardinia.^[246][247] True prawns and polychelids appeared during the Triassic, while true crabs and caridean shrimps emerged in the Jurassic.^{[248][249][250][251][252][253]} A well-preserved Jurassic example is the decapod Aeger spinipes from the Solnhofen limestone of Germany.^[254] Marine lobsters and crabs underwent an evolutionary radiation in the Cretaceous, potentially driven by the co-evolutionary radiation of predatory teleost fishes.^[251][255] Extinct fossil groups of uncertain phylogenetic placement include the Thylacocephala (Silurian to Cretaceous), possessing a large bivalved carapace and raptorial appendages, and the enigmatic Cyclida (Carboniferous to Cretaceous).^[256] No fossil record is currently known for krill, mystacocarids, or cephalocarids.^{[257][258][259][260]}

Human Interactions and Economic Importance

Crustaceans represent a major component of global marine fisheries and aquaculture.^{[261][262][263][264]} Over 7.9 million tonnes of crustaceans were harvested annually by the mid-2000s, expanding to 10.7 million tonnes in 2007.^{[261][262][265]} In 2016, global aquaculture alone generated 7.86 million tonnes of crustaceans, with marine shrimps and prawns accounting for 5.18 million tonnes.^[266] Over 60 percent by weight of all crustaceans caught for consumption consists of shrimp and prawns, and nearly 80 percent of worldwide production is based in Asia, with China producing more than half of the global total.^[262][266] Popular edible species include the European lobster (Homarus gammarus), the American lobster (Homarus americanus), the red king crab (Paralithodes camtschaticus), the brown crab (Cancer pagurus), the gazami crab (Portunus trituberculatus), and penaeid prawns like Litopenaeus vannamei and Penaeus monodon.^{[263][266][267][268][269]} Non-decapod consumption is limited, though over 118,000 tonnes of Antarctic krill are harvested annually for animal feed, dietary supplements, and biochemical extraction.^[262][270]

In aquatic ecosystems and water purification, planktonic microcrustaceans act as natural biofilters, consuming bacteria, microalgae, and organic detritus in municipal reservoirs.^[129] Live Artemia salina cysts, nauplii, and Daphnia are cultivated globally as feed for fish fry in commercial hatcheries and domestic aquaria.^[36][271] Several crustacean species serve as laboratory model organisms in developmental biology, genetics, and ecotoxicology, notably the amphipod Parhyale hawaiensis and the water flea Daphnia pulex.^[98][272] Chitin and chitosan extracted from waste crab and shrimp carapaces are utilized in pharmaceutical, agricultural, and industrial chemical applications.^[36]

Conversely, crustaceans can impose significant economic and medical burdens.^{[25][36][273][274][275]} Sessile acorn barnacles settle in dense colonies on marine vessels, subsea installations, and intake pipes, generating heavy biofouling that increases hydrodynamic drag by up to 35 percent and necessitates expensive hull careening and anti-fouling treatments.^{[25][36][274]} Wood-boring isopods, notably Limnoria lignorum (the gribble), and amphipods like Chelura terebrans bore into submerged timber pilings, bridges, and wooden docks, causing structural degradation.^[25][36] Parasitic species such as fish lice (Branchiura) and cymothoid isopods can cause large losses in fish ponds and fish farms during mass outbreaks.^[25][275] Several freshwater crustaceans act as intermediate hosts for human parasites: copepods transmit the broad fish tapeworm (Dibothriocephalus latus) and the Guinea worm (Dracunculus medinensis), while freshwater crabs (such as Potamon) transmit the human lung fluke (Paragonimus).^{[36][273][276][277]} Tongue worms (Pentastomida) cause visceral or nasopharyngeal pentastomiasis (halzoun syndrome) in humans when raw nymph-infected tissues are consumed.^[278][279] Crustaceans can also cause food allergies in humans.^[280]

Studies of pain in crustaceans have reached differing conclusions: a study on prawns by one team found no acid receptors on their antennae.^[281] According to Robert Elwood of Queen's University Belfast, research increasingly suggests that crustaceans can perceive suffering, and in a study he co-authored, acid placed on the antennae of prawns (Palaemon elegans) caused them to groom the affected area for more than five minutes, which could be a protective response to pain.^[282][283] Since 1 March 2018, Switzerland has banned plunging live lobsters into boiling water, requiring that they first be stunned by electric shock or by mechanical destruction of the brain to spare them unnecessary suffering, and the philosopher Jonathan Birch has proposed applying the precautionary principle so that the whole order of decapods is covered by animal protection laws requiring that animals be killed quickly with minimal suffering.^[284][285]

References

1. Calman, William Thomas (1911). "Crustacea" . In Chisholm, Hugh (ed.). Encyclopædia Britannica. Vol. 7 (11th ed.). Cambridge University Press. p. 552.
2. Rota-Stabelli, Omar; Kayal, Ehsan; Gleeson, Dianne; et al. (2010). "Ecdysozoan Mitogenomics: Evidence for a Common Origin of the Legged Invertebrates, the Panarthropoda". Genome Biology and Evolution. 2: 425–440. doi:10.1093/gbe/evq030. PMC 2998192. PMID 20624745.
3. Brusca, Richard C.; Silveira, Fábio Lang da (2007). Invertebrados 2. ed. Rio de Janeiro: Guanabara Koogan. ISBN 852771258X. OCLC 124028674
4. Brusca, Moore a Giribet 2023, s. 730–731. (BRUSCA, R. C.; MOORE, Wendy; GIRIBET, G, 2023. Invertebrates. 4. vyd. New York: Oxford University Press. ISBN 9780197554418. S. 731. (anglicky))
5. Charbonnier (2025), p. 8.
6. Lipke B. Holthuis (1991). "Introduction". Marine Lobsters of the World. FAO Species Catalogue, Volume 13. Food and Agriculture Organization. pp. 1–2. ISBN 978-92-5-103027-1.
7. FOREST, Jacques; VON VAUPEL KLEIN, J. C., 2012. Treatise on zoology. Volume 3, The crustacea : anatomy, taxonomy, biology. Leiden: Brill. S. 403–405.
8. http://www.archive.org/details/mthbrnnichiizo00br
9. Arnaud Zucker, Aristote et les classifications zoologiques, Éditions Peeters, 2005 (ISBN 90-429-1660-5, lire en ligne), p. 145
10. (en) Rose Gulledge, « “GALLERY OF CARCINOLOGISTS” Returns to IZ-NMNH », sur No Bones, Smithsonian Institute, 15 novembre 2016.
11. Informations lexicographiques et étymologiques de « carcinologie » dans le Trésor de la langue française informatisé, sur le site du Centre national de ressources textuelles et lexicales
12. "Cephalon". Crustacean Glossary. Natural History Museum of Los Angeles County. Archived from the original on 2011-07-27. Retrieved 2016-09-10.
13. Schram a Koenemann 2022, s. 4–5. (SCHRAM, Frederick R.; KOENEMANN, Stefan, 2022. Evolution and Phylogeny of Pancrustacea: A Story of Scientific Method. 1. vyd. New York: Oxford University Press. Dostupné online. ISBN 978-0-19-536576-4, ISBN 978-0-19-752185-4. doi:10.1093/oso/9780195365764.001.0001. S. 53. (anglicky))
14. Шарова, 2002, с. 349. (Шарова И. Х. Зоология беспозвоночных. — М.: Владос, 2002. — 592 с. — ISBN 5-691-00332-1.)
15. Inga Mohrbeck, Pedro Martínez Arbizu & Thomas Glatzel (October 2010). "Tantulocarida (Crustacea) from the Southern Ocean deep sea, and the description of three new species of Tantulacus Huys, Andersen & Kristensen, 1992". Systematic Parasitology. 77 (2): 131–151. doi:10.1007/s11230-010-9260-0. PMID 20852984. S2CID 7325858.
16. HUTCHINS, Michael; CRAIG, S. F.; THONEY, Dennis A.; SCHLAGER, Neil, 2003. Protostomes. 2. vyd. Farmington Hills, MI: Gale. (Grzimek's Animal Life Encyclopedia; sv. 2). S. 198. (anglicky)
17. McClain C. R., Boyer A. G. Biodiversity and body size are linked across metazoans (англ.) // Proceedings of the Royal Society B: Biological Sciences : журнал. — 2009. — Vol. 296, no. 1665. — P. 2209—2215. — doi:10.1098/rspb.2009.0245. — PMID 19324730. Архивировано 3 марта 2016 года. (Дата обращения: 2 марта 2015)
18. Brusca, Moore a Giribet 2023, s. 659–662. (BRUSCA, R. C.; MOORE, Wendy; GIRIBET, G, 2023. Invertebrates. 4. vyd. New York: Oxford University Press. ISBN 9780197554418. S. 731. (anglicky))
19. Brusca, Moore a Giribet 2023, s. 699–701. (BRUSCA, R. C.; MOORE, Wendy; GIRIBET, G, 2023. Invertebrates. 4. vyd. New York: Oxford University Press. ISBN 9780197554418. S. 731. (anglicky))
20. Armengol, J. et al. (1986): Artròpodes (I). Història Natural dels Països Catalans, 9. Barcelona: Enciclopèdia Catalana, S. A. ISBN 84-85194-84-5
21. "Thorax". Crustacean Glossary. Natural History Museum of Los Angeles County. Archived from the original on 2011-07-27. Retrieved 2016-09-10.
22. "Abdomen". Crustacean Glossary. Natural History Museum of Los Angeles County. Archived from the original on 2011-07-27. Retrieved 2016-09-10.
23. Brusca, R. C. & Brusca, G. J., 2005. Invertebrados, 2.ª edición. McGraw-Hill-Interamericana, Madrid (etc.), XXVI+1005 pp. ISBN 0-87893-097-3.
24. Ракообразные // Большая советская энциклопедия : [в 30 т.] / гл. ред. А. М. Прохоров. — 3-е изд. — М. : Советская энциклопедия, 1969—1978.
25. Krzysztof Jażdżewski: Podtyp: skorupiaki – Crustacea. W: Zoologia: Stawonogi. Szczękoczułkopodobne, skorupiaki. T. 2, cz. 1. Czesław Błaszak (red. nauk.). Warszawa: Wydawnictwo Naukowe PWN, 2011. ISBN 978-83-01-16568-0.
26. "Cephalothorax". Crustacean Glossary. Natural History Museum of Los Angeles County. Archived from the original on 2011-07-27. Retrieved 2016-09-10.
27. Шарова, 2002, с. 349—351. (Шарова И. Х. Зоология беспозвоночных. — М.: Владос, 2002. — 592 с. — ISBN 5-691-00332-1.)
28. "Carapace". Crustacean Glossary. Natural History Museum of Los Angeles County. Archived from the original on 2011-07-27. Retrieved 2016-09-10.
29. Шарова, 2002, с. 352. (Шарова И. Х. Зоология беспозвоночных. — М.: Владос, 2002. — 592 с. — ISBN 5-691-00332-1.)
30. MOTYČKA, Vladimír; ROLLER, Zdeněk, 2001. Bezobratlí (1). Praha: Albatros. (Svět zvířat; sv. 10). S. 102.
31. SMRŽ, Jaroslav, 2015. Základy biologie, ekologie a systému bezobratlých živočichů. Praha: Karolinum Press. ISBN 8024622580, ISBN 9788024622583. S. 128.
32. P. J. Hayward; J. S. Ryland (1995). Handbook of the marine fauna of north-west Europe. Oxford University Press. ISBN 978-0-19-854055-7. Retrieved 2016-09-10.
33. Motyčka a Roller 2001, s. 93. (MOTYČKA, Vladimír; ROLLER, Zdeněk, 2001. Bezobratlí (1). Praha: Albatros. (Svět zvířat; sv. 10). S. 102.)
34. Догель, 1981, с. 296—297. (Догель В. А. Зоология беспозвоночных. — 7-е изд.. — М.: Высшая школа, 1981. — 614 с.)
35. "Antennule". Crustacean Glossary. Natural History Museum of Los Angeles County. Archived from the original on 2013-11-05. Retrieved 2016-09-10.
36. Integrated Taxonomic Information System (ITIS), www.itis.gov, CC0 https://doi.org/10.5066/F7KH0KBK, consulté le 30 novembre 2015.
37. Brusca, Richard (2018). Invertebrados. Rio de Janeiro: Gen - Grupo Editorial Nacional - Editora Guanabara Koogan LTDA
38. Thorp a Rogers 2015, s. 673–675. (THORP, James H.; ROGERS, D. Christopher, 2015. In: Ecology and General Biology (Thorp and Covich’s Freshwater Invertebrates – Volume I). 4. vyd. London: Academic Press. ISBN 978-0-12-385027-0, ISBN 0-12-385027-4. S. 710. (anglicky))
39. Castro-Huber, Marine Biology, fjerde utgave, McGraw Hill 2003, side 135.
40. Brusca, Richard C. (2005). Invertebrados. [S.l.]: McGraw Hill. ISBN 9788448602468. OCLC 1085749697
41. Догель, 1981, с. 295. (Догель В. А. Зоология беспозвоночных. — 7-е изд.. — М.: Высшая школа, 1981. — 614 с.)
42. Шарова, 2002, с. 351. (Шарова И. Х. Зоология беспозвоночных. — М.: Владос, 2002. — 592 с. — ISBN 5-691-00332-1.)
43. Fritsch, Martin; Richter, Stefan (September 5, 2022). "How body patterning might have worked in the evolution of arthropods—A case study of the mystacocarid Derocheilocaris remanei (Crustacea, Oligostraca)". Journal of Experimental Zoology Part B: Molecular and Developmental Evolution. 338 (6): 342–359. Bibcode:2022JEZB..338..342F. doi:10.1002/jez.b.23140. PMID 35486026. S2CID 248430846.
44. Burkenroad, M. D. (1963). "The evolution of the Eucarida (Crustacea, Eumalacostraca), in relation to the fossil record". Tulane Studies in Geology. 2 (1): 1–17.
45. Brusca, Moore a Giribet 2023, s. 724–725. (BRUSCA, R. C.; MOORE, Wendy; GIRIBET, G, 2023. Invertebrates. 4. vyd. New York: Oxford University Press. ISBN 9780197554418. S. 731. (anglicky))
46. "Telson". Crustacean Glossary. Natural History Museum of Los Angeles County. Archived from the original on 2011-07-27. Retrieved 2016-09-10.
47. Догель, 1981, с. 308—309. (Догель В. А. Зоология беспозвоночных. — 7-е изд.. — М.: Высшая школа, 1981. — 614 с.)
48. Akira Sakurai. "Closed and Open Circulatory System". Georgia State University. Archived from the original on 2016-09-17. Retrieved 2016-09-10.
49. Brusca, Moore a Giribet 2023, s. 717–719. (BRUSCA, R. C.; MOORE, Wendy; GIRIBET, G, 2023. Invertebrates. 4. vyd. New York: Oxford University Press. ISBN 9780197554418. S. 731. (anglicky))
50. Догель, 1981, с. 300. (Догель В. А. Зоология беспозвоночных. — 7-е изд.. — М.: Высшая школа, 1981. — 614 с.)
51. Thorp a Rogers 2015, s. 675. (THORP, James H.; ROGERS, D. Christopher, 2015. In: Ecology and General Biology (Thorp and Covich’s Freshwater Invertebrates – Volume I). 4. vyd. London: Academic Press. ISBN 978-0-12-385027-0, ISBN 0-12-385027-4. S. 710. (anglicky))
52. Sokolowicz, Carolina (2010). «História Natural de Benthana caireles (Isopoda: Oniscidea)» (PDF). Universidade federal do Rio grande do Sul. Consultado em 24 de junho de 2019. Cópia arquivada (PDF) em 7 de novembro de 2025
53. WATLING, Les; THIEL, Martin, 2013. Functional morphology and diversity. New York: Oxford University Press. (The natural history of the Crustacea; sv. 1). ISBN 978-0-19-539803-8. S. 378.
54. Klaus Urich (1994). "Respiratory pigments". Comparative Animal Biochemistry. Springer. pp. 249–287. ISBN 978-3-540-57420-0.
55. Urich K. Respiratory pigments // Comparative Animal Biochemistry (англ.) / K. Urich; Translated from the German by P. J. King. — Berlin—Heidelberg: Springer-Verlag, 1994. — P. 249—287. — 782 p. — ISBN 3-540-57420-4. (Дата обращения: 31 января 2017)
56. Sedlák 2002, s. 84. (SEDLÁK, Edmund, 2002. Zoologie bezobratlých. Brno: Přírodovědecká fakulta MU. ISBN 80-210-2892-0. S. 90.)
57. MAYNARD, DONALD M. (1960). «CIRCULATION AND HEART FUNCTION». Elsevier: 161–226. ISBN 9780123956286
58. H. J. Ceccaldi. Anatomy and physiology of digestive tract of Crustaceans Decapods reared in aquaculture (PDF). AQUACOP, IFREMER. Actes de Colloque 9. pp. 243–259.
59. Brusca, Moore a Giribet 2023, s. 714–717. (BRUSCA, R. C.; MOORE, Wendy; GIRIBET, G, 2023. Invertebrates. 4. vyd. New York: Oxford University Press. ISBN 9780197554418. S. 731. (anglicky))
60. Шарова, 2002, с. 353. (Шарова И. Х. Зоология беспозвоночных. — М.: Владос, 2002. — 592 с. — ISBN 5-691-00332-1.)
61. RUPPERT; FOX; BARNES;, E.E.; R. S.; R. D. (2005). Zoologia dos Invertebrados 7ª ed. São Paulo: Rocca
62. Watling a Thiel 2013, s. 251–255. (WATLING, Les; THIEL, Martin, 2013. Functional morphology and diversity. New York: Oxford University Press. (The natural history of the Crustacea; sv. 1). ISBN 978-0-19-539803-8. S. 378.)
63. Шарова, 2002, с. 352—353. (Шарова И. Х. Зоология беспозвоночных. — М.: Владос, 2002. — 592 с. — ISBN 5-691-00332-1.)
64. Догель, 1981, с. 298. (Догель В. А. Зоология беспозвоночных. — 7-е изд.. — М.: Высшая школа, 1981. — 614 с.)
65. Догель, 1981, с. 299. (Догель В. А. Зоология беспозвоночных. — 7-е изд.. — М.: Высшая школа, 1981. — 614 с.)
66. Watling a Thiel 2013, s. 399. (WATLING, Les; THIEL, Martin, 2013. Functional morphology and diversity. New York: Oxford University Press. (The natural history of the Crustacea; sv. 1). ISBN 978-0-19-539803-8. S. 378.)
67. Smrž 2015, s. 129–130. (SMRŽ, Jaroslav, 2015. Základy biologie, ekologie a systému bezobratlých živočichů. Praha: Karolinum Press. ISBN 8024622580, ISBN 9788024622583. S. 128.)
68. Watling a Thiel 2013, s. 404–406. (WATLING, Les; THIEL, Martin, 2013. Functional morphology and diversity. New York: Oxford University Press. (The natural history of the Crustacea; sv. 1). ISBN 978-0-19-539803-8. S. 378.)
69. Ghiselin, Michael T. (2005). "Crustacean". Encarta. Microsoft.
70. Brusca, Moore a Giribet 2023, s. 719–720. (BRUSCA, R. C.; MOORE, Wendy; GIRIBET, G, 2023. Invertebrates. 4. vyd. New York: Oxford University Press. ISBN 9780197554418. S. 731. (anglicky))
71. Догель, 1981, с. 306. (Догель В. А. Зоология беспозвоночных. — 7-е изд.. — М.: Высшая школа, 1981. — 614 с.)
72. Шарова, 2002, с. 355—356. (Шарова И. Х. Зоология беспозвоночных. — М.: Владос, 2002. — 592 с. — ISBN 5-691-00332-1.)
73. Alessandro Poli e Elena Fabbri, Fisiologia degli animali marini, 2ª ed., EdiSES, 2018, ISBN 978-88-7959-8569.
74. Thorp a Rogers 2015, s. 675–676. (THORP, James H.; ROGERS, D. Christopher, 2015. In: Ecology and General Biology (Thorp and Covich’s Freshwater Invertebrates – Volume I). 4. vyd. London: Academic Press. ISBN 978-0-12-385027-0, ISBN 0-12-385027-4. S. 710. (anglicky))
75. Brusca, Moore a Giribet 2023, s. 720–724. (BRUSCA, R. C.; MOORE, Wendy; GIRIBET, G, 2023. Invertebrates. 4. vyd. New York: Oxford University Press. ISBN 9780197554418. S. 731. (anglicky))
76. Brusca, R.; Brusca, G. J. (2007). Invertebrados 2.ª ed. Rio de Janeiro: Guanabara Koogan. 968 páginas
77. Догель, 1981, с. 301. (Догель В. А. Зоология беспозвоночных. — 7-е изд.. — М.: Высшая школа, 1981. — 614 с.)
78. Giribet a Edgecombe 2020, s. 245–246. (GIRIBET, Gonzalo; EDGECOMBE, Gregory, 2020. The Invertebrate Tree of Life. Princeton, NJ Oxford: Princeton University Press. ISBN 978-0-691-19706-7, ISBN 0-691-19706-7. OCLC 1129197548 S. 234. (anglicky))
79. Шарова, 2002, с. 356. (Шарова И. Х. Зоология беспозвоночных. — М.: Владос, 2002. — 592 с. — ISBN 5-691-00332-1.)
80. Догель, 1981, с. 303. (Догель В. А. Зоология беспозвоночных. — 7-е изд.. — М.: Высшая школа, 1981. — 614 с.)
81. Догель, 1981, с. 302. (Догель В. А. Зоология беспозвоночных. — 7-е изд.. — М.: Высшая школа, 1981. — 614 с.)
82. Шарова, 2002, с. 357. (Шарова И. Х. Зоология беспозвоночных. — М.: Владос, 2002. — 592 с. — ISBN 5-691-00332-1.)
83. Шарова, 2002, с. 357—358. (Шарова И. Х. Зоология беспозвоночных. — М.: Владос, 2002. — 592 с. — ISBN 5-691-00332-1.)
84. Nielsen 2012, s. 247. (NIELSEN, Claus, 2012. Animal Evolution : Interrelationships of The Living Phyla. 3. vyd. Oxford: OUP. ISBN 978-0-19-162530-5, ISBN 0-19-162530-2. S. 244. (anglicky))
85. Ruppert, Edward E. (1996). Zoologia dos invertebrados 6. ed. São Paulo: Roca. ISBN 8572411496. OCLC 940019119
86. Шарова, 2002, с. 358. (Шарова И. Х. Зоология беспозвоночных. — М.: Владос, 2002. — 592 с. — ISBN 5-691-00332-1.)
87. MARSHALL, Justin; CRONIN, Thomas W.; KLEINLOGEL, Sonja. Stomatopod eye structure and function: A review. Arthropod Structure & Development. 2007-12, roč. 36, čís. 4, s. 420–448. Dostupné online [cit. 2026-07-18]. doi:10.1016/j.asd.2007.01.006. (anglicky)
88. MELLON, JR., DeForest, 2014. Sensory systems of crustaceans. In: DERBY, Charles Dorsett; THIEL, Martin. Nervous systems and control of behavior. New York: Oxford University Press. ISBN 978-0-19-979171-2. S. 49–52.
89. Motyčka a Roller 2001, s. 113. (MOTYČKA, Vladimír; ROLLER, Zdeněk, 2001. Bezobratlí (1). Praha: Albatros. (Svět zvířat; sv. 10). S. 102.)
90. Mellon, Jr. 2014, s. 65–66. (MELLON, JR., DeForest, 2014. Sensory systems of crustaceans. In: DERBY, Charles Dorsett; THIEL, Martin. Nervous systems and control of behavior. New York: Oxford University Press. ISBN 978-0-19-979171-2. S. 49–52.)
91. Moyes, C. D.:Schulte, P. M. (ed.). «Principios de Fisiologia Animal». Porto Alegre: Artmed, 2010, 2 ed. Consultado em 9 de junho de 2019. Cópia arquivada em 23 de novembro de 2024
92. Saxena, A. (ed.). «The Book of Crustacea». New Delhi: DPH, 2005. Consultado em 9 de junho de 2019. Cópia arquivada em 23 de janeiro de 2025
93. LIU, Lei; TAO, Dandan; WANG, Chunlin. Autotomy and Regeneration of Appendages in Crustaceans: A Review. Journal of Ocean University of China. 2024-06-01, roč. 23, čís. 3, s. 731–742. Dostupné online [cit. 2026-07-30]. ISSN 1993-5021. doi:10.1007/s11802-024-5747-1. (anglicky)
94. Догель, 1981, с. 311. (Догель В. А. Зоология беспозвоночных. — 7-е изд.. — М.: Высшая школа, 1981. — 614 с.)
95. Pechenik, J. A. (ed.). «Biologia dos Invertebrados». Porto Alegre: Artmed, 2016. 7 ed. Consultado em 2 de junho de 2019. Cópia arquivada em 23 de janeiro de 2025
96. Carlisle, D. B.; Knowles, F. (ed.). «Endocrine Control In Crustaceans». London: Cambridge, 1959. Consultado em 2 de junho de 2019. Cópia arquivada em 23 de novembro de 2024
97. Watling, L.; Thiel, M. (ed.). «Phisiology: The Natural History of Crustacea». New York: Oxford, 2015. v. 4. Consultado em 6 de junho de 2019. Cópia arquivada em 23 de novembro de 2024
98. Derby, Charles (Charles Dorsett), editor. Thiel, Martin, 1962- editor. Nervous systems and control of behavior. [S.l.: s.n.] ISBN 9780199791736. OCLC 889675086
99. Wiese, K. (ed.). «The Crustacean Nervous System». Berlin: Springer, 2002. Consultado em 1 de junho de 2019. Cópia arquivada em 23 de janeiro de 2025
100. "Crustacean (arthropod)". Encyclopædia Britannica. 5 May 2023.
101. Cothran a Thiel 2020, s. 145–148. (COTHRAN, Rickey D.; THIEL, Martin, 2020. Reproductive biology. New York: Oxford University Press. (The natural history of the Crustacea; sv. 6). ISBN 978-0-19-068855-4. S. 61.)
102. Crustacean (arthropod) // Encyclopædia Britannica. — [Архивировано 10 ноября 2013 года.]
103. G. L. Pesce. "Remipedia Yager, 1981".
104. D. E. Aiken; V. Tunnicliffe; C. T. Shih; L. D. Delorme. "Crustacean". The Canadian Encyclopedia. Archived from the original on 2011-06-07. Retrieved 2016-09-10.
105. NEUFELD, Christopher J.; PALMER, A. Richard. Precisely proportioned: intertidal barnacles alter penis form to suit coastal wave action. Proceedings. Biological Sciences. 2008-05-07, roč. 275, čís. 1638, s. 1081–1087. Dostupné online [cit. 2026-07-28]. ISSN 0962-8452. doi:10.1098/rspb.2007.1760. PMID 18252665.
106. COTHRAN, Rickey D.; THIEL, Martin, 2020. Reproductive biology. New York: Oxford University Press. (The natural history of the Crustacea; sv. 6). ISBN 978-0-19-068855-4. S. 61.
107. Schram a Koenemann 2022, s. 127. (SCHRAM, Frederick R.; KOENEMANN, Stefan, 2022. Evolution and Phylogeny of Pancrustacea: A Story of Scientific Method. 1. vyd. New York: Oxford University Press. Dostupné online. ISBN 978-0-19-536576-4, ISBN 978-0-19-752185-4. doi:10.1093/oso/9780195365764.001.0001. S. 53. (anglicky))
108. Smrž 2015, s. 125. (SMRŽ, Jaroslav, 2015. Základy biologie, ekologie a systému bezobratlých živočichů. Praha: Karolinum Press. ISBN 8024622580, ISBN 9788024622583. S. 128.)
109. Lopes, Diogo (Fevereiro de 2012). «Produção do camarão-rosa Farfantepenaeus brasiliensis em sistemas com mínima renovação de água» (PDF). Universidade Federal do Rio Grande. Consultado em 23 de junho de 2019. Cópia arquivada (PDF) em 23 de junho de 2019
110. Шарова, 2002, с. 358—359. (Шарова И. Х. Зоология беспозвоночных. — М.: Владос, 2002. — 592 с. — ISBN 5-691-00332-1.)
111. Cothran a Thiel 2020, s. 127. (COTHRAN, Rickey D.; THIEL, Martin, 2020. Reproductive biology. New York: Oxford University Press. (The natural history of the Crustacea; sv. 6). ISBN 978-0-19-068855-4. S. 61.)
112. Alan P. Covich; James H. Thorp (2001). "Introduction to the Subphylum Crustacea". In James H. Thorp; Alan P. Covich (eds.). Ecology and classification of North American freshwater invertebrates (2nd ed.). Academic Press. pp. 777–798. ISBN 978-0-12-690647-9. Retrieved 2016-09-10.
113. Alan P. Covich & James H. Thorp. Introduction to the Subphylum Crustacea // Ecology and classification of North American freshwater invertebrates (англ.) / James H. Thorp & Alan P. Covich. — 2nd. — Academic Press, 2001. — P. 777—798. — ISBN 978-0-12-690647-9.
114. Thorp a Rogers 2015, s. 697. (THORP, James H.; ROGERS, D. Christopher, 2015. In: Ecology and General Biology (Thorp and Covich’s Freshwater Invertebrates – Volume I). 4. vyd. London: Academic Press. ISBN 978-0-12-385027-0, ISBN 0-12-385027-4. S. 710. (anglicky))
115. Догель, 1981, с. 313. (Догель В. А. Зоология беспозвоночных. — 7-е изд.. — М.: Высшая школа, 1981. — 614 с.)
116. Brusca, Moore a Giribet 2023, s. 727–730. (BRUSCA, R. C.; MOORE, Wendy; GIRIBET, G, 2023. Invertebrates. 4. vyd. New York: Oxford University Press. ISBN 9780197554418. S. 731. (anglicky))
117. Шарова, 2002, с. 360. (Шарова И. Х. Зоология беспозвоночных. — М.: Владос, 2002. — 592 с. — ISBN 5-691-00332-1.)
118. "Zoea". Oxford English Dictionary (online ed.). Oxford University Press. (Subscription or participating institution membership required.)
119. Schram a Koenemann 2022, s. 36–38. (SCHRAM, Frederick R.; KOENEMANN, Stefan, 2022. Evolution and Phylogeny of Pancrustacea: A Story of Scientific Method. 1. vyd. New York: Oxford University Press. Dostupné online. ISBN 978-0-19-536576-4, ISBN 978-0-19-752185-4. doi:10.1093/oso/9780195365764.001.0001. S. 53. (anglicky))
120. Schram a Koenemann 2022, s. 43. (SCHRAM, Frederick R.; KOENEMANN, Stefan, 2022. Evolution and Phylogeny of Pancrustacea: A Story of Scientific Method. 1. vyd. New York: Oxford University Press. Dostupné online. ISBN 978-0-19-536576-4, ISBN 978-0-19-752185-4. doi:10.1093/oso/9780195365764.001.0001. S. 53. (anglicky))
121. W. F. R. Weldon (July 1889). "Note on the function of the spines of the Crustacean zoœa" (PDF). Journal of the Marine Biological Association of the United Kingdom. 1 (2): 169–172. Bibcode:1889JMBUK...1..169W. doi:10.1017/S0025315400057994. S2CID 54759780. Archived from the original (PDF) on 2011-07-17.
122. Шарова, 2002, с. 360—361. (Шарова И. Х. Зоология беспозвоночных. — М.: Владос, 2002. — 592 с. — ISBN 5-691-00332-1.)
123. Thorp a Rogers 2015, s. 678–679. (THORP, James H.; ROGERS, D. Christopher, 2015. In: Ecology and General Biology (Thorp and Covich’s Freshwater Invertebrates – Volume I). 4. vyd. London: Academic Press. ISBN 978-0-12-385027-0, ISBN 0-12-385027-4. S. 710. (anglicky))
124. Догель, 1981, с. 310. (Догель В. А. Зоология беспозвоночных. — 7-е изд.. — М.: Высшая школа, 1981. — 614 с.)
125. Srivastava, Shikha; Dahal, Sumedha; Naidu, Sharanya J.; Anand, Deepika; Gopalakrishnan, Vidya; Kooloth Valappil, Rajendran; Raghavan, Sathees C. (24 January 2017). "DNA double-strand break repair in Penaeus monodon is predominantly dependent on homologous recombination". DNA Research. 24 (2): 117–128. doi:10.1093/dnares/dsw059. PMC 5397610. PMID 28431013.
126. Rhee, J. S.; Kim, B. M.; Choi, B. S.; Lee, J. S. (2012). "Expression pattern analysis of DNA repair-related and DNA damage response genes revealed by 55K oligomicroarray upon UV-B irradiation in the intertidal copepod, Tigriopus japonicus". Comparative Biochemistry and Physiology. Toxicology & Pharmacology. 155 (2): 359–368. doi:10.1016/j.cbpc.2011.10.005. PMID 22051804.
127. Brusca, R.C. & Brusca, G. J, 2007. Invertebrados. Segunda edição. Editora Guanabara-Koogan, Rio de Janeiro. 968 pp.
128. Thorp a Rogers 2015, s. 681. (THORP, James H.; ROGERS, D. Christopher, 2015. In: Ecology and General Biology (Thorp and Covich’s Freshwater Invertebrates – Volume I). 4. vyd. London: Academic Press. ISBN 978-0-12-385027-0, ISBN 0-12-385027-4. S. 710. (anglicky))
129. Brusca, Moore a Giribet 2023, s. 708–711. (BRUSCA, R. C.; MOORE, Wendy; GIRIBET, G, 2023. Invertebrates. 4. vyd. New York: Oxford University Press. ISBN 9780197554418. S. 731. (anglicky))
130. Narchi, Walter (1973). Crustáceos. São Paulo, SP: Editora da Universidade de São Paulo
131. Southward, A.J. 1955. Feeding of barnacles. Nature 175:1124–1125
132. Crisp, D.J., and A.J. Southward. 1961. Different types of cirral activity of barnacles. Philosophical Transactions of the Royal Society 243:271–307.
133. Friedman, M.M., and J.R. Strickler. 1975. Chemoreceptors and feeding in calanoid copepods (Arthropoda: Crustacea). Proceedings of the Natural Academy of Sciences USA 72:4185–4188
134. Hamner, W.M. 1988. Biomechanics of filter feeding in the Antarctic krill Euphausia superba: review of past work and new observations. Journal of Crustacean Biology 8:149–163
135. Nicol, E.A.T. 1932. The feeding habits of the Galatheidea. Journal of Marine Biology Association of the U.K. 18:87–106.
136. Trager, G.C., D. Coughlin, A. Genin, Y. Achituv, and A. Gangopadhyay. 1992. Foraging to the rhythm of ocean waves: porcelain crabs and barnacles synchronize motions with flow oscillations. Journal of Experimental Marine Biology and Ecology 164:73–86
137. Vogel, F. 1984. Comparative and functional morphology of the spoon-tipped setae on the second maxillipeds in Dotilla Stimpson, 1858 (Decapoda, Brachyura, Ocypodidae). Crustaceana 47:225–234.
138. Fielder, D.R. 1970. The feeding behaviour of the sand crab Scopimera inflata (Decapoda, Ocypodidae). Journal of Zoology, London 160:35–49.
139. Griffis, R.B., and T.H. Suchanek. 1991. A model of burrow architecture and trophic modes in thalassinidean shrimp (Decapoda: Thalassinidea). Marine Ecology Progress Series 79:171–183.
140. Taylor, G.M. 2000. Maximum force production: why are crabs so strong? Proceedings of the Royal Society of London B: Biological Sciences 267:1475–1480.
141. Lau, C.J. 1987. Feeding behavior of the Hawaiian slipper lobster, Scyllarides squammosus, with a review of decapod feeding tactics on molluscan prey. Bulletin of Marine Science 41:378–391.
142. Hutchins et al. 2003, s. 200. (HUTCHINS, Michael; CRAIG, S. F.; THONEY, Dennis A.; SCHLAGER, Neil, 2003. Protostomes. 2. vyd. Farmington Hills, MI: Gale. (Grzimek's Animal Life Encyclopedia; sv. 2). S. 198. (anglicky))
143. Caldwell, R.L., and M.J. Childress. 1990. Prey selection and processing in a stomatopod crustacean. Pages 143–164 in R.N. Hughes, editor. Behavioural mechanisms of food selection (2nd edition). Springer, Berlin, Germany.
144. Patek, S.N., W.L. Korff, and R.L. Caldwell. 2004. Deadly strike mechanisms of a mantis shrimp. Nature 428:819–820.
145. Hutchins et al. 2003, s. 170. (HUTCHINS, Michael; CRAIG, S. F.; THONEY, Dennis A.; SCHLAGER, Neil, 2003. Protostomes. 2. vyd. Farmington Hills, MI: Gale. (Grzimek's Animal Life Encyclopedia; sv. 2). S. 198. (anglicky))
146. Schram a Koenemann 2022, s. 735. (SCHRAM, Frederick R.; KOENEMANN, Stefan, 2022. Evolution and Phylogeny of Pancrustacea: A Story of Scientific Method. 1. vyd. New York: Oxford University Press. Dostupné online. ISBN 978-0-19-536576-4, ISBN 978-0-19-752185-4. doi:10.1093/oso/9780195365764.001.0001. S. 53. (anglicky))
147. Matt Soniak, Meet the World’s Only Known Venomous Crustacean, <www.mentalfloss.com, läst 2016-05-22
148. Thurston, M.H. 1979. Scavenging abyssal amphipods from the North-East Atlantic Ocean. Marine Biology 51:55–68.
149. Shulenberger, E., and R.R. Hessler. 1974. Scavenging abyssal benthic amphipods trapped under oligotrophic central North Pacific Gyre waters. Marine Biology 28:185–187.
150. Bui, T. H. Hanh; Lee, Shing Yip (1 de setembro de 2015). «Potential contributions of gut microbiota to the nutrition of the detritivorous sesarmid crab Parasesarma erythodactyla». Marine Biology. 162 (10): 1969–1981. ISSN 0025-3162. doi:10.1007/s00227-015-2723-8
151. MARIN, Ivan N.; TIUNOV, Alexei V. Terrestrial crustaceans (Arthropoda, Crustacea): taxonomic diversity, terrestrial adaptations, and ecological functions. ZooKeys. 2023-07-13, roč. 1169, s. 95–162. Dostupné online [cit. 2026-07-10]. ISSN 1313-2970. doi:10.3897/zookeys.1169.97812. PMID 38328027.
152. M. Zimmer et al.: Cellulose digestion and phenol oxidation in coastal isopods (Crustacea: Isopoda). In: Marine Biology, Band 140, Nr. 6, 2002, S. 1207–1213, doi:10.1007/s00227-002-0800-2.
153. Benjamin J. Allardyce, Stuart M. Linton: Purification and characterisation of endo-β-1, 4-glucanase and laminarinase enzymes from the gecarcinid land crab Gecarcoidea natalis and the aquatic crayfish Cherax destructor. In: Journal of Experimental Biology, Band 211, Nr. 14, 2008, S. 2275–2287.
154. VOLF, Petr; HORÁK, Petr, 2007. Paraziti a jejich biologie. Praha: Triton. ISBN 9788073870089, ISBN 8073870088. S. 255.
155. Шарова, 2002, с. 348. (Шарова И. Х. Зоология беспозвоночных. — М.: Владос, 2002. — 592 с. — ISBN 5-691-00332-1.)
156. Schram a Koenemann 2022, s. 79. (SCHRAM, Frederick R.; KOENEMANN, Stefan, 2022. Evolution and Phylogeny of Pancrustacea: A Story of Scientific Method. 1. vyd. New York: Oxford University Press. Dostupné online. ISBN 978-0-19-536576-4, ISBN 978-0-19-752185-4. doi:10.1093/oso/9780195365764.001.0001. S. 53. (anglicky))
157. Volf a Horák 2007, s. 256. (VOLF, Petr; HORÁK, Petr, 2007. Paraziti a jejich biologie. Praha: Triton. ISBN 9788073870089, ISBN 8073870088. S. 255.)
158. Schram a Koenemann 2022, s. 112. (SCHRAM, Frederick R.; KOENEMANN, Stefan, 2022. Evolution and Phylogeny of Pancrustacea: A Story of Scientific Method. 1. vyd. New York: Oxford University Press. Dostupné online. ISBN 978-0-19-536576-4, ISBN 978-0-19-752185-4. doi:10.1093/oso/9780195365764.001.0001. S. 53. (anglicky))
159. Volf a Horák 2007, s. 257–258. (VOLF, Petr; HORÁK, Petr, 2007. Paraziti a jejich biologie. Praha: Triton. ISBN 9788073870089, ISBN 8073870088. S. 255.)
160. Догель, 1981, с. 317—319. (Догель В. А. Зоология беспозвоночных. — 7-е изд.. — М.: Высшая школа, 1981. — 614 с.)
161. Brusca, Moore a Giribet 2023, s. 693–694. (BRUSCA, R. C.; MOORE, Wendy; GIRIBET, G, 2023. Invertebrates. 4. vyd. New York: Oxford University Press. ISBN 9780197554418. S. 731. (anglicky))
162. Корнев П. Н. Первое нахождение представителей подкласса Tantulocarida в Белом море // Зоология беспозвоночных : журнал. — 2004. — Т. 1, № 1. Архивировано 29 октября 2013 года. (PDF)
163. MEHLHORN, Heinz. Encyclopedia of Parasitology. Berlin: Springer, 2008. ISBN 978-3-540-48994-8. S. 299–300. (anglicky)
164. "Crabs, lobsters, prawns and other crustaceans". Australian Museum. January 5, 2010. Retrieved 2016-09-10.
165. Brusca, Moore a Giribet 2023, s. 714. (BRUSCA, R. C.; MOORE, Wendy; GIRIBET, G, 2023. Invertebrates. 4. vyd. New York: Oxford University Press. ISBN 9780197554418. S. 731. (anglicky))
166. VAUGHAN, David Brendan; GRUTTER, Alexandra Sara; COSTELLO, Mark John. Cleaner fishes and shrimp diversity and a re‐evaluation of cleaning symbioses. Fish and Fisheries. 2017-07, roč. 18, čís. 4, s. 698–716. Dostupné online [cit. 2026-08-07]. ISSN 1467-2960. doi:10.1111/faf.12198. (anglicky)
167. MIKÁT, Michael. Eusocialita. In: ŠPINKA, Marek; HAVLÍČEK, Jan; ŠTOLHOFEROVÁ, Iveta. Etologie – Mechanismy, ontogeneze, funkce a evoluce chování živočichů. 1. vyd. Praha: Academia, 2024. ISBN 978-80-200-3556-1. S. 482.
168. PENG, Xiaotong; DU, Mengran; GEBRUK, Andrey. Flourishing chemosynthetic life at the greatest depths of hadal trenches. Nature. 2025-09, roč. 645, čís. 8081, s. 679–685. Dostupné online [cit. 2026-08-05]. ISSN 1476-4687. doi:10.1038/s41586-025-09317-z. (anglicky)
169. Hutchins et al. 2003, s. 300. (HUTCHINS, Michael; CRAIG, S. F.; THONEY, Dennis A.; SCHLAGER, Neil, 2003. Protostomes. 2. vyd. Farmington Hills, MI: Gale. (Grzimek's Animal Life Encyclopedia; sv. 2). S. 198. (anglicky))
170. SEDLÁČEK, František. Chování v prostoru a času, chování a orientace v prostoru. In: ŠPINKA, Marek; HAVLÍČEK, Jan; ŠTOLHOFEROVÁ, Iveta. Etologie – Mechanismy, ontogeneze, funkce a evoluce chování živočichů. 1. vyd. Praha: Academia, 2024. ISBN 978-80-200-3556-1. S. 160.
171. Virtue, P. D.; Nichols, P. D.; Nicols, S. (1997). "Dietary-related mechanisms of survival in Euphasia superba: biochemical changes during long term starvation and bacteria as a possible source of nutrition.". In Bruno Battaglia; Valencia, José; Walton, D. W. H. (eds.). Antarctic communities: species, structure, and survival. Cambridge University Press. ISBN 978-0-521-48033-8. Retrieved 2016-09-10.
172. ATTENBOROUGH, David; BUTFIELD, Colin. Oceán: poslední divočina na Zemi (původním názvem: Ocean: Earth's Last Wilderness). Překlad Jitka Jeníková. Praha: Práh, 2025. 392 s. ISBN 978-80-7696-072-5. S. 300–303.
173. Thorp a Rogers 2015, s. 679–680. (THORP, James H.; ROGERS, D. Christopher, 2015. In: Ecology and General Biology (Thorp and Covich’s Freshwater Invertebrates – Volume I). 4. vyd. London: Academic Press. ISBN 978-0-12-385027-0, ISBN 0-12-385027-4. S. 710. (anglicky))
174. Догель, 1981, с. 312. (Догель В. А. Зоология беспозвоночных. — 7-е изд.. — М.: Высшая школа, 1981. — 614 с.)
175. ŠTĚRBA, Otakar. Pramen života. Praha: Panorama, 1986. S. 16–20.
176. Hutchins et al. 2003, s. 245. (HUTCHINS, Michael; CRAIG, S. F.; THONEY, Dennis A.; SCHLAGER, Neil, 2003. Protostomes. 2. vyd. Farmington Hills, MI: Gale. (Grzimek's Animal Life Encyclopedia; sv. 2). S. 198. (anglicky))
177. Hutchins et al. 2003, s. 125. (HUTCHINS, Michael; CRAIG, S. F.; THONEY, Dennis A.; SCHLAGER, Neil, 2003. Protostomes. 2. vyd. Farmington Hills, MI: Gale. (Grzimek's Animal Life Encyclopedia; sv. 2). S. 198. (anglicky))
178. Шарова, 2002, с. 383. (Шарова И. Х. Зоология беспозвоночных. — М.: Владос, 2002. — 592 с. — ISBN 5-691-00332-1.)
179. Brusca, Moore a Giribet 2023, s. 703–708. (BRUSCA, R. C.; MOORE, Wendy; GIRIBET, G, 2023. Invertebrates. 4. vyd. New York: Oxford University Press. ISBN 9780197554418. S. 731. (anglicky))
180. Gollasch, Stephan (October 30, 2006). "Eriocheir sinensis" (PDF). Global Invasive Species Database. Invasive Species Specialist Group. doi:10.15468/ybwd3x. Archived from the original (PDF) on 2017-12-24. Retrieved 2016-09-10.
181. John J. McDermott (1999). "The western Pacific brachyuran Hemigrapsus sanguineus (Grapsidae) in its new habitat along the Atlantic coast of the United States: feeding, cheliped morphology and growth". In Schram, Frederick R.; Klein, J. C. von Vaupel (eds.). Crustaceans and the biodiversity crisis: Proceedings of the Fourth International Crustacean Congress, Amsterdam, the Netherlands, July 20–24, 1998. Koninklijke Brill. pp. 425–444. ISBN 978-90-04-11387-9. Retrieved 2016-09-10.
182. PILLAY, Deena. Ecosystem Engineering by Thalassinidean Crustaceans: Response Variability, Contextual Dependencies and Perspectives on Future Research. Diversity. 2019-04-19, roč. 11, čís. 4, s. 64. Dostupné online [cit. 2026-08-08]. ISSN 1424-2818. doi:10.3390/d11040064. (anglicky)
183. HÄNFLING, Bernd; EDWARDS, François; GHERARDI, Francesca. Invasive alien Crustacea: dispersal, establishment, impact and control. BioControl. 2011-08, roč. 56, čís. 4, s. 573–595. Dostupné online [cit. 2026-08-08]. ISSN 1386-6141. doi:10.1007/s10526-011-9380-8. (anglicky)
184. J. B. Shurin & J. E. Havel, (en) « Hydrologic connections and overland dispersal in an exotic freshwater crustacean » in : BiologicalInvasions n°4, pp. 431–439, 2002.
185. Galil, Bella; Froglia, Carlo; Noël, Pierre (2002). Briand, Frederic (ed.). CIESM Atlas of Exotic Species in the Mediterranean: Vol 2 Crustaceans. Paris, Monaco: CIESM Publishers. p. 192. ISBN 92-990003-2-8.
186. ZRZAVÝ, Jan. Fylogeneze živočišné říše. 1. vyd. Praha: Scientia 255 s. ISBN 80-86960-08-0, ISBN 978-80-86960-08-1. OCLC 124086610 S. 161–163.
187. Czesław Jura: Bezkręgowce. Podstawy morfologii funkcjonalne, systematyki i filogenezy. Wyd. 3. Warszawa: PWN, 2005, s. 423–434,469, 476.
188. Joel W. Martin; George E. Davis (2001). An Updated Classification of the Recent Crustacea (PDF). Natural History Museum of Los Angeles County. pp. 1–132. Archived from the original (PDF) on 2013-05-12. Retrieved 2009-12-14.
189. Huys, Rony (2003). "An Updated Classification of the Recent Crustacea". review. Journal of Crustacean Biology. 23 (2): 495–497. Bibcode:2003JCBio..23..495H. doi:10.1163/20021975-99990355.
190. Martin, J.W. & Davis, G.E. 2001. An updated classification of the recent Crustacea. Natural History Museum of Los Angeles County, Science Series 39 : 132p.
191. Kurt Schminke: Crustacea, Krebse, Seite 565 in Wilfried Westheide & Reinhard Rieger (Hrsg., 2007): Spezielle Zoologie - Teil 1: Einzeller und Wirbellose Tiere (2. Aufl.). Elsevier, Spektrum Akademischer Verlag, München. ISBN 3-8274-1575-6
192. Hynek Burda, Gero Hilken, Jan Zrzavý: Systematische Zoologie. UTB, Stuttgart; : 1. Aufl. 2008, Seite 187-188, ISBN 3-8252-3119-4
193. Jerome C. Regier; Jeffrey W. Shultz; Andreas Zwick; April Hussey; Bernard Ball; Regina Wetzer; Joel W. Martin; Clifford W. Cunningham (February 25, 2010). "Arthropod relationships revealed by phylogenomic analysis of nuclear protein-coding sequences". Nature. 463 (7284): 1079–1083. Bibcode:2010Natur.463.1079R. doi:10.1038/nature08742. PMID 20147900. S2CID 4427443.
194. Björn M. von Reumont; Ronald A. Jenner; Matthew A. Wills; Emiliano Dell'Ampio; Günther Pass; Ingo Ebersberger; Benjamin Meyer; Stefan Koenemann; Thomas M. Iliffe; Alexandros Stamatakis; Oliver Niehuis; Karen Meusemann; Bernhard Misof (March 2012). "Pancrustacean phylogeny in the light of new phylogenomic data: support for Remipedia as the possible sister group of Hexapoda". Molecular Biology and Evolution. 29 (3): 1031–1045. doi:10.1093/molbev/msr270. PMID 22049065.
195. Oakley, Todd H.; Wolfe, Joanna M.; Lindgren, Annie R.; Zaharoff, Alexander K. (January 2013). "Phylotranscriptomics to bring the understudied into the fold: monophyletic ostracoda, fossil placement, and pancrustacean phylogeny". Molecular Biology and Evolution. 30 (1): 215–233. doi:10.1093/molbev/mss216. PMID 22977117.
196. BRUSCA, R. C.; MOORE, Wendy; GIRIBET, G, 2023. Invertebrates. 4. vyd. New York: Oxford University Press. ISBN 9780197554418. S. 731. (anglicky)
197. GIRIBET, Gonzalo; EDGECOMBE, Gregory, 2020. The Invertebrate Tree of Life. Princeton, NJ Oxford: Princeton University Press. ISBN 978-0-691-19706-7, ISBN 0-691-19706-7. OCLC 1129197548 S. 234. (anglicky)
198. Vargas, Pablo (Vargas Gómez); Zardoya, Rafael (2012). El árbol de la vida : sistemática y evolución de los seres vivos. Madrid: [s.n.] ISBN 9788461597406. OCLC 830350033
199. Regier, Jerome C.; Shultz, Jeffrey W.; Zwick, Andreas; Hussey, April; Ball, Bernard; Wetzer, Regina; Martin, Joel W.; Cunningham, Clifford W. (fevereiro de 2010). «Arthropod relationships revealed by phylogenomic analysis of nuclear protein-coding sequences». Nature. 463 (7284): 1079–1083. ISSN 0028-0836. doi:10.1038/nature08742
200. Richter, S (2002). «The Tetraconata concept: hexapod-crustacean relationships and the phylogeny of Crustacea». Organisms Diversity & Evolution (em inglês). 2 (3): 217–237. doi:10.1078/1439-6092-00048. Cópia arquivada em 23 de janeiro de 2025
201. Todd H. Oakley, Joanna M. Wolfe, Annie R. Lindgren, Alexander K. Zaharoff: Phylotranscriptomics to Bring the Understudied into the Fold: Monophyletic Ostracoda, Fossil Placement, and Pancrustacean Phylogeny. In: Molecular Biology and Evolution. Band 30, Nr. 1, 2013, S. 215–233, doi:10.1093/molbev/mss216.
202. ZRZAVÝ, J.; ŠTYS, P. The basic body plan of arthropods: insights from evolutionary morphology and developmental biology. Journal of Evolutionary Biology. 1997-05, roč. 10, čís. 3, s. 353–367. Dostupné online [cit. 2026-07-09]. ISSN 1010-061X. doi:10.1046/j.1420-9101.1997.10030353.x. (anglicky)
203. DOHLE, W. Are the insects terrestrial crustaceans? A discussion of some new facts and arguments and the proposal of the proper name 'Tetraconata' for the monophyletic unit Crustacea + Hexapoda. Ann. Soc. Entomol. Fr.. 2001, roč. 37, čís. 1–2, s. 85–103.
204. Schram a Koenemann 2022, s. 54–55. (SCHRAM, Frederick R.; KOENEMANN, Stefan, 2022. Evolution and Phylogeny of Pancrustacea: A Story of Scientific Method. 1. vyd. New York: Oxford University Press. Dostupné online. ISBN 978-0-19-536576-4, ISBN 978-0-19-752185-4. doi:10.1093/oso/9780195365764.001.0001. S. 53. (anglicky))
205. G. Giribet; G. D. Edgecombe, W.C. Wheeler. Arthropod phylogeny based on eight molecular loci and morphology. „Nature”. 413, s. 157–161, 2001.
206. Brusca, Richard C. (2016). Invertebrates (3rd ed.). Sunderland, MA: Sinauer Associates. p. 222. ISBN 978-1-60535-375-3.
207. «WoRMS - World Register of Marine Species». www.marinespecies.org. Consultado em 26 de junho de 2019. Cópia arquivada em 8 de junho de 2026
208. Crustacea. In: World Register of Marine Species. WoRMS Editorial Board, abgerufen am 17. August 2026.
209. Multicrustacea. In: World Register of Marine Species. WoRMS Editorial Board, abgerufen am 17. August 2026.
210. Schwentner, M; Combosch, DJ; Nelson, JP; Giribet, G (2017). "A Phylogenomic Solution to the Origin of Insects by Resolving Crustacean-Hexapod Relationships". Current Biology. 27 (12): 1818–1824.e5. Bibcode:2017CBio...27E1818S. doi:10.1016/j.cub.2017.05.040. PMID 28602656.
211. Jerome C. Regier; Jeffrey W. Shultz; Andreas Zwick; April Hussey; Bernard Ball; Regina Wetzer; Joel W. Martin; Clifford W. Cunningham. Arthropod relationships revealed by phylogenomic analysis of nuclear protein-coding sequences. „Nature”. 463 (7284), s. 1079–1083, 2010. DOI: 10.1038/nature08742.
212. Oligostraca. In: World Register of Marine Species. WoRMS Editorial Board, abgerufen am 17. August 2026.
213. J. Zrzavý, V. Hypsa, M. Vlásková: Arthropod phylogeny: taxonomic congruence, total evidence and conditional combination approaches to morphological and molecular data sets. W: Arthropod Relationships. Richard A. Fortey, Richard H. Thomas (red.). London: Chapman & Hall, 1998, s. 97–107.
214. Jesus Lozano-Fernandez, Mattia Giacomelli, James F. Fleming, Albert Chen, Jakob Vinther, Philip Francis Thomsen, Henrik Glenner, Ferran Palero, David A. Legg, Thomas M. Iliffe, Davide Pisani, Jørgen Olesen. Pancrustacean Evolution Illuminated by Taxon-Rich Genomic-Scale Data Sets with an Expanded Remipede Sampling. „Genome Biol. Evol.”. 1 (8), s. 2055–2070, 2019. DOI: 10.1093/gbe/evz097.
215. BERNOT, James P; OWEN, Christopher L; WOLFE, Joanna M, 2023. Major revisions in pancrustacean phylogeny and evidence of sensitivity to taxon sampling. Molecular Biology and Evolution. Roč. 40, čís. 8, s. msad175. Dostupné online [cit. 2026-07-09]. ISSN 1537-1719. doi:10.1093/molbev/msad175.
216. YU, Hiu Yan; CHU, Ka Hou; TSANG, Ling Ming. Incomplete lineage sorting and long-branch attraction confound phylogenomic inference of Pancrustacea. Frontiers in Ecology and Evolution. 2024-01-30, roč. 12. Dostupné online [cit. 2026-07-09]. ISSN 2296-701X. doi:10.3389/fevo.2024.1243221.
217. James P. Bernot, Christopher L. Owen, Joanna M. Wolfe, Kenneth Meland, Jørgen Olesen, Keith A. Crandall: Major Revisions in Pancrustacean Phylogeny and Evidence of Sensitivity to Taxon Sampling. In: Molecular Biology and Evolution. Band 40, Nr. 8, 2023, doi:10.1093/molbev/msad175.
218. Allotriocarida. In: World Register of Marine Species. WoRMS Editorial Board, abgerufen am 17. August 2026.
219. Martin Schwentner, David J. Combosch, Joey Pakes Nelson, Gonzalo Giribet. A Phylogenomic Solution to the Origin of Insects by Resolving Crustacean-Hexapod Relationships. „Current Biology”. 27, s. 1–7, 2017. Elsevier. DOI: 10.1016/j.cub.2017.05.040.
220. SCHWENTNER, Martin; COMBOSCH, David J.; PAKES NELSON, Joey. A phylogenomic Solution to the origin of insects by resolving crustacean-hexapod relationships. Current Biology. 2017-06, roč. 27, čís. 12, s. 1818–1824.e5. Dostupné online [cit. 2026-07-09]. doi:10.1016/j.cub.2017.05.040. (anglicky)
221. VON REUMONT, Björn M.; BURMESTER, Thorsten. Remipedia and the evolution of hexapods. In: Encyclopedia of life sciences. Chichester: John Wiley & Sons, Ltd, 2010. Dostupné online. ISBN 978-0-470-01590-2.
222. Giribet, G.; Edgecombe, G.D. (2020). The Invertebrate Tree of Life. Princeton University Press. p. 21. ISBN 978-0-6911-7025-1. Retrieved 27 May 2023.
223. Hutchins et al. 2003, s. 132. (HUTCHINS, Michael; CRAIG, S. F.; THONEY, Dennis A.; SCHLAGER, Neil, 2003. Protostomes. 2. vyd. Farmington Hills, MI: Gale. (Grzimek's Animal Life Encyclopedia; sv. 2). S. 198. (anglicky))
224. Motyčka a Roller 2001, s. 100. (MOTYČKA, Vladimír; ROLLER, Zdeněk, 2001. Bezobratlí (1). Praha: Albatros. (Svět zvířat; sv. 10). S. 102.)
225. Schram a Koenemann 2022, s. 152. (SCHRAM, Frederick R.; KOENEMANN, Stefan, 2022. Evolution and Phylogeny of Pancrustacea: A Story of Scientific Method. 1. vyd. New York: Oxford University Press. Dostupné online. ISBN 978-0-19-536576-4, ISBN 978-0-19-752185-4. doi:10.1093/oso/9780195365764.001.0001. S. 53. (anglicky))
226. Корнев П. Н., Чесунов А. В. Тантулокариды — микроскопические обитатели Белого моря (рус.) // Природа : журнал. — Наука, 2005. — № 2. Архивировано 12 января 2012 года. (PDF)
227. Olney, Matthew. "Ostracods". An insight into micropalaeontology. University College, London. Retrieved 2016-09-10.
228. BRUSCA; MOORE; SHUSTER, Richard C.; Wendy; Stephen M. (2018). Invertebrados 3ª ed. Rio de Janeiro: Guanabara Koogan. 900 páginas
229. Brusca, Moore a Giribet 2023, s. 732–734. (BRUSCA, R. C.; MOORE, Wendy; GIRIBET, G, 2023. Invertebrates. 4. vyd. New York: Oxford University Press. ISBN 9780197554418. S. 731. (anglicky))
230. Шарова, 2002, с. 363. (Шарова И. Х. Зоология беспозвоночных. — М.: Владос, 2002. — 592 с. — ISBN 5-691-00332-1.)
231. Maas, Andreas; Waloszek, Dieter (1 de março de 2005). «Phosphatocopina – ostracode-like sister group of Eucrustacea». Hydrobiologia (em inglês). 538 (1): 139–152. ISSN 1573-5117. doi:10.1007/PL00021866
232. BRUSCA; MOORE; SHUSTER, Richard C.; Wendy; Stephen M. (2018). Invertebrados 3ª ed. Rio de Janeiro: Guanabara Koogan. pp. 900 e 901
233. http://palaeo.gly.bris.ac.uk/Palaeofiles/Fossilgroups/Crustacea/fossils.html Archyvuota kopija 2016-09-07 iš Wayback Machine projekto.
234. Giribet a Edgecombe 2020, s. 257. (GIRIBET, Gonzalo; EDGECOMBE, Gregory, 2020. The Invertebrate Tree of Life. Princeton, NJ Oxford: Princeton University Press. ISBN 978-0-691-19706-7, ISBN 0-691-19706-7. OCLC 1129197548 S. 234. (anglicky))
235. DE OLIVEIRA ALMEIDA, Waltécio et al. Morphological support for the phylogenetic positioning of Pentastomida and related fossils. Biotemas, v. 21, n. 3, p. 81-90, 2008.
236. Hutchins et al. 2003, s. 312. (HUTCHINS, Michael; CRAIG, S. F.; THONEY, Dennis A.; SCHLAGER, Neil, 2003. Protostomes. 2. vyd. Farmington Hills, MI: Gale. (Grzimek's Animal Life Encyclopedia; sv. 2). S. 198. (anglicky))
237. Paul Selden & John R. Nudds (2004). "The Rhynie Chert". Evolution of Fossil Ecosystems (2nd ed.). Manson Publishing. pp. 47–58. ISBN 978-1-84076-041-5.
238. Wills, M. A. (1998). Fortey, R. A.; Thomas, R. H., eds. «A phylogeny of recent and fossil Crustacea derived from morphological characters». Dordrecht: Springer Netherlands. The Systematics Association Special Volume Series (em inglês): 189–209. ISBN 9789401149044. doi:10.1007/978-94-011-4904-4_15.pdf
239. Koenemann, Stefan; Schram, Frederick R.; Hönemann, Mario; Iliffe, Thomas M. (12 April 2007). "Phylogenetic analysis of Remipedia (Crustacea)". Organisms Diversity & Evolution. 7 (1): 33–51. Bibcode:2007ODivE...7...33K. doi:10.1016/j.ode.2006.07.001.
240. "Fossil Record". Fossil Groups: Crustacea. University of Bristol. Archived from the original on 2016-09-07. Retrieved 2016-09-10.
241. Koenemann, Stefan; Iliffe, Thomas M.; Rust, Jes; Bergmann, Alexandra; Stemme, Torben; Hartke, Tamara R.; Neiber, Marco T. (19 de maio de 2011). «Global Biodiversity and Phylogenetic Evaluation of Remipedia (Crustacea)». PLOS ONE (em inglês). 6 (5): e19627. ISSN 1932-6203. doi:10.1371/journal.pone.0019627. Cópia arquivada em 19 de outubro de 2025
242. Schram, Frederick; Hof, Cees H. J.; Mapes, Royal H. & Snowdon, Polly (2003). "Paleozoic cumaceans (Crustacea, Malacostraca, Peracarida) from North America". Contributions to Zoology. 72 (1): 1–16. doi:10.1163/18759866-07201001.
243. Schram, Frederick R. (August 28, 1970). "Isopod from the Pennsylvanian of Illinois". Science. 169 (3948): 854–855. Bibcode:1970Sci...169..854S. doi:10.1126/science.169.3948.854. PMID 5432581. S2CID 31851291.
244. Hof, Cees H. J. (1998). "Fossil stomatopods (Crustacea: Malacostraca) and their phylogenetic impact". Journal of Natural History. 32 (10 & 11): 1567–1576. Bibcode:1998JNatH..32.1567H. doi:10.1080/00222939800771101.
245. Haug, Joachim T.; Haug, Carolin; Maas, Andreas; Kutschera, Verena; Waloszek, Dieter (21 de setembro de 2010). «Evolution of mantis shrimps (Stomatopoda, Malacostraca) in the light of new Mesozoic fossils». BMC Evolutionary Biology. 10 (1). 290 páginas. ISSN 1471-2148. doi:10.1186/1471-2148-10-290
246. Baucon, A.; Ronchi, A.; Felletti, F.; Neto de Carvalho, C. (2014). "Evolution of Crustaceans at the edge of the end-Permian crisis: ichnonetwork analysis of the fluvial succession of Nurra (Permian-Triassic, Sardinia, Italy)". Palaeogeography, Palaeoclimatology, Palaeoecology. 410: 74. Bibcode:2014PPP...410...74B. doi:10.1016/j.palaeo.2014.05.034. Archived from the original on July 6, 2022. Retrieved May 19, 2022.
247. Baucon, Andrea; Ronchi, Ausonio; Felletti, Fabrizio; Neto de Carvalho, Carlos (setembro de 2014). «Evolution of Crustaceans at the edge of the end-Permian crisis: Ichnonetwork analysis of the fluvial succession of Nurra (Permian–Triassic, Sardinia, Italy)». Palaeogeography, Palaeoclimatology, Palaeoecology (em inglês). 410: 74–103. doi:10.1016/j.palaeo.2014.05.034. Cópia arquivada em 20 de fevereiro de 2025
248. Crean, Robert P. D. (November 14, 2004). "Dendrobranchiata". Order Decapoda. University of Bristol. Archived from the original on February 29, 2012. Retrieved February 25, 2010.
249. Karasawa, Hiroaki; Takahashi, Fumio; Doi, Eiji; Ishida, Hideo (2003). "First notice of the family Coleiidae Van Straelen (Crustacea: Decapoda: Eryonoides) from the upper Triassic of Japan". Paleontological Research. 7 (4): 357–362. Bibcode:2003PalRe...7..357K. doi:10.2517/prpsj.7.357. S2CID 129330859.
250. Chace, Fenner A. Jr.; Manning, Raymond B. (1972). "Two new caridean shrimps, one representing a new family, from marine pools on Ascension Island (Crustacea: Decapoda: Natantia)". Smithsonian Contributions to Zoology. 131 (131): 1–18. doi:10.5479/si.00810282.131. S2CID 53067015.
251. Wägele, J. W. (December 1989). "On the influence of fishes on the evolution of benthic crustaceans". Zeitschrift für Zoologische Systematik und Evolutionsforschung. 27 (4): 297–309. doi:10.1111/j.1439-0469.1989.tb00352.x.
252. http://palaeo.gly.bris.ac.uk/Palaeofiles/Fossilgroups/Decapoda/dendrobranchiata.html Archyvuota kopija 2012-02-29 iš Wayback Machine projekto.
253. http://si-pddr.si.edu/dspace/bitstream/10088/5471/2/SCtZ-0131-Lo_res.pdf
254. «Um verdadeiro parque jurássico». Ciência Hoje. Consultado em 23 de junho de 2019. Cópia arquivada em 16 de março de 2026
255. Tshudy, Dale; Donaldson, W. Steven; Collom, Christopher; et al. (2005). "Hoploparia albertaensis, a new species of clawed lobster (Nephropidae) from the Late Coniacean, shallow-marine Bad Heart Formation of northwestern Alberta, Canada". Journal of Paleontology. 79 (5): 961–968. doi:10.1666/0022-3360(2005)079[0961:HAANSO]2.0.CO;2. S2CID 131067067.
256. HEGNA, Thomas A.; LUQUE, Javier; WOLFE, Joanna M. The Fossil Record of the Pancrustacea. In: POORE, Gary C. B.; THIEL, Martin. Evolution and biogeography. New York: Oxford University Press, 2020. Dostupné online. ISBN 978-0-19-063784-2.
257. Hessler, R. R. (1984). "Cephalocarida: living fossil without a fossil record". In N. Eldredge; S. M. Stanley (eds.). Living Fossils. New York: Springer Verlag. pp. 181–186. ISBN 978-3-540-90957-6.
258. "Antarctic Prehistory". Australian Antarctic Division. July 29, 2008. Archived from the original on September 30, 2009. Retrieved February 25, 2010.
259. Meurant, Gerard (28 de setembro de 1982). The Biology of Crustacea: Volume 1: Systematics, The Fossil Record, And Biogeography (em inglês). [S.l.]: Academic Press. ISBN 9780323139250. Cópia arquivada em 23 de novembro de 2024
260. Hessler, Robert R. (1984). Eldredge, Niles; Stanley, Steven M., eds. «Cephalocarida: Living Fossil Without a Fossil Record». New York, NY: Springer New York. Casebooks in Earth Sciences (em inglês): 181–186. ISBN 9781461382713. doi:10.1007/978-1-4613-8271-3_20
261. "The State of World Fisheries and Aquaculture 2018 – Meeting the sustainable development goals" (PDF). fao.org. Rome: Food and Agriculture Organization of the United Nations. 2018.
262. "FIGIS: Global Production Statistics 1950–2007". Food and Agriculture Organization. Retrieved 2016-09-10.
263. Hutchins et al. 2003, s. 201. (HUTCHINS, Michael; CRAIG, S. F.; THONEY, Dennis A.; SCHLAGER, Neil, 2003. Protostomes. 2. vyd. Farmington Hills, MI: Gale. (Grzimek's Animal Life Encyclopedia; sv. 2). S. 198. (anglicky))
264. Hudinaga, M. 1942. Reproduction, development and rearing of Penaeus japonicus. Jap.J. Zool. 10 : 305-393
265. Global Production Statistics 1950-2017. Food and Agriculture Organization of United Nations. Fisheris and Aquaculture Department. [dostęp 2020-01-12].
266. Albert G.J. Tacon. Global Trends in Aquaculture and Compound Aquafeed Production. „World Aquaculture”, s. 33–46, 2018.
267. Michał Grabowski: Nadrząd: Eucarida. W: Zoologia: Stawonogi. Szczękoczułkopodobne, skorupiaki. T. 2, cz. 1. Czesław Błaszak (red. nauk.). Warszawa: Wydawnictwo Naukowe PWN, 2011, s. 328–344. ISBN 978-83-01-16568-0.
268. http://www.fao.org/fishery/culturedspecies/Penaeus_monodon/en
269. http://www.fao.org/fishery/culturedspecies/Litopenaeus_vannamei/en
270. Hutchins et al. 2003, s. 187. (HUTCHINS, Michael; CRAIG, S. F.; THONEY, Dennis A.; SCHLAGER, Neil, 2003. Protostomes. 2. vyd. Farmington Hills, MI: Gale. (Grzimek's Animal Life Encyclopedia; sv. 2). S. 198. (anglicky))
271. Hutchins et al. 2003, s. 137. (HUTCHINS, Michael; CRAIG, S. F.; THONEY, Dennis A.; SCHLAGER, Neil, 2003. Protostomes. 2. vyd. Farmington Hills, MI: Gale. (Grzimek's Animal Life Encyclopedia; sv. 2). S. 198. (anglicky))
272. PARIS, Mathilde; WOLFF, Carsten; PATEL, Nipam H. The crustacean model Parhyale hawaiensis. Current Topics in Developmental Biology. 2022, roč. 147, s. 199–230. Dostupné online [cit. 2026-08-09]. ISSN 1557-8933. doi:10.1016/bs.ctdb.2022.02.001. PMID 35337450.
273. Volf a Horák 2007, s. 165. (VOLF, Petr; HORÁK, Petr, 2007. Paraziti a jejich biologie. Praha: Triton. ISBN 9788073870089, ISBN 8073870088. S. 255.)
274. Hutchins et al. 2003, s. 276. (HUTCHINS, Michael; CRAIG, S. F.; THONEY, Dennis A.; SCHLAGER, Neil, 2003. Protostomes. 2. vyd. Farmington Hills, MI: Gale. (Grzimek's Animal Life Encyclopedia; sv. 2). S. 198. (anglicky))
275. Hutchins et al. 2003, s. 290. (HUTCHINS, Michael; CRAIG, S. F.; THONEY, Dennis A.; SCHLAGER, Neil, 2003. Protostomes. 2. vyd. Farmington Hills, MI: Gale. (Grzimek's Animal Life Encyclopedia; sv. 2). S. 198. (anglicky))
276. Volf a Horák 2007, s. 223. (VOLF, Petr; HORÁK, Petr, 2007. Paraziti a jejich biologie. Praha: Triton. ISBN 9788073870089, ISBN 8073870088. S. 255.)
277. Volf a Horák 2007, s. 183. (VOLF, Petr; HORÁK, Petr, 2007. Paraziti a jejich biologie. Praha: Triton. ISBN 9788073870089, ISBN 8073870088. S. 255.)
278. Volf a Horák 2007, s. 258. (VOLF, Petr; HORÁK, Petr, 2007. Paraziti a jejich biologie. Praha: Triton. ISBN 9788073870089, ISBN 8073870088. S. 255.)
279. Michał Grabowski: Gromada: wrzęchy – Pentastomida. W: Zoologia: Stawonogi. Szczękoczułkopodobne, skorupiaki. T. 2, cz. 1. Czesław Błaszak (red. nauk.). Warszawa: Wydawnictwo Naukowe PWN, 2011, s. 277–278. ISBN 978-83-01-16568-0.
280. Äyriäiset ja nilviäiset (muun muassa simpukat, mustekalat, osterit ja etanat) Ruokavirasto. Viitattu 10.1.2021.
281. (en) Sakshi Puri et Zen Faulkes, « Do Decapod Crustaceans Have Nociceptors for Extreme pH? », PLOS ONE, vol. 5, no 4,‎ 20 avril 2010, e10244 (ISSN 1932-6203, PMID 20422026, PMCID PMC2857684, DOI 10.1371/journal.pone.0010244, lire en ligne, consulté le 12 décembre 2020)
282. (en-GB) Emine Saner, « Is it wrong to boil lobsters alive? », The Guardian,‎ 11 février 2018 (ISSN 0261-3077, lire en ligne, consulté le 26 mars 2019)
283. (en) Stuart Barr, Peter R. Laming, Jaimie T. A. Dick et Robert W. Elwood, « Nociception or pain in a decapod crustacean? », Animal Behaviour, vol. 75, no 3,‎ 1er mars 2008, p. 745–751 (ISSN 0003-3472, DOI 10.1016/j.anbehav.2007.07.004, lire en ligne, consulté le 12 décembre 2020)
284. « Souffrance animale : pourquoi la Suisse interdit de plonger les homards dans l'eau bouillante », sur L'Obs, Le Nouvel Obs, 11 janvier 2018 (consulté le 25 mars 2019).
285. Birch, Jonathan (2017) Animal sentience and the precautionary principle. Animal Sentience 16(1): https://www.wellbeingintlstudiesrepository.org/animsent/vol2/iss16/1/